| [1] |
Fraley RT, Rogers SG, Horsch RB, Sanders PR, Flick JS, et al. 1983. Expression of bacterial genes in plant cells. |
| [2] |
Herrera-Estrella L, Depicker A, Van Montagu M, Schell J. 1983. Expression of chimaeric genes transferred into plant cells using a Ti-plasmid-derived vector. |
| [3] |
Zhou GY, Weng J, Zeng Y, Huang J, Qian S, et al. 1983. Introduction of exogenous DNA into cotton embryos. |
| [4] |
Mohammed S, Samad AA, Rahmat Z. 2019. Agrobacterium-mediated transformation of rice: constraints and possible solutions. |
| [5] |
Steinberger AR, Voytas DF. 2025. Virus-induced gene editing free from tissue culture. |
| [6] |
Gordon-Kamm WJ, Spencer TM, Mangano ML, Adams TR, Daines RJ, et al. 1990. Transformation of maize cells and regeneration of rertile transgenic plants. |
| [7] |
Wu M, Chen A, Li X, Li X, Hou X, et al. 2024. Advancements in delivery strategies and non-tissue culture regeneration systems for plant genetic transformation. |
| [8] |
Morikawa H, Yamada Y. 1985. Capillary Microinjection into Protoplasts and Intranuclear Localization of Injected Materials. |
| [9] |
Paszkowski J, Shillito RD, Saul M, Mandák V, Hohn T, et al. 1984. Direct gene transfer to plants. |
| [10] |
Wang P, Si H, Li C, Xu Z, Guo H, et al. 2025. Plant genetic transformation: achievements, current status and future prospects. |
| [11] |
Kaeppler HF, Somers DA, Rines HW, Cockburn AF. 1992. Silicon carbide fiber-mediated stable transformation of plant cells. |
| [12] |
Zhao X, Meng Z, Wang Y, Chen W, Sun C, et al. 2017. Pollen magnetofection for genetic modification with magnetic nanoparticles as gene carriers. |
| [13] |
Su W, Xu M, Radani Y, Yang L. 2023. Technological development and application of plant genetic transformation. |
| [14] |
Cao X, Xie H, Song M, Lu J, Ma P, et al. 2023. Cut−dip−budding delivery system enables genetic modifications in plants without tissue culture. |
| [15] |
Tamizi AA, Md-Yusof AA, Mohd-Zim NA, Nazaruddin NH, Sekeli R, et al. 2023. Agrobacterium-mediated in planta transformation of cut coleoptile: a new, simplified, and tissue culture-independent method to deliver the CRISPR/Cas9 system in rice. |
| [16] |
Zhong H, Li C, Yu W, Zhou HP, Lieber T, et al. 2024. A fast and genotype-independent in planta Agrobacterium-mediated transformation method for soybean. |
| [17] |
Lowe K, Wu E, Wang N, Hoerster G, Hastings C, et al. 2016. Morphogenic regulators Baby boom and Wuschel improve monocot transformation. |
| [18] |
Lowe K, La Rota M, Hoerster G, Hastings C, Wang N, et al. 2018. Rapid genotype “independent” Zea mays L. (maize) transformation via direct somatic embryogenesis. |
| [19] |
Wang N, Arling M, Hoerster G, Ryan L, Wu E, et al. 2020. An efficient gene excision system in maize. |
| [20] |
Che P, Wu E, Simon MK, Anand A, Lowe K, et al. 2022. Wuschel2 enables highly efficient CRISPR/Cas-targeted genome editing during rapid de novo shoot regeneration in sorghum. |
| [21] |
Chen J, Tomes S, Gleave AP, Hall W, Luo Z, et al. 2022. Significant improvement of apple (Malus domestica Borkh.) transgenic plant production by pre-transformation with a Baby boom transcription factor. |
| [22] |
Iwase A, Mita K, Nonaka S, Ikeuchi M, Koizuka C, et al. 2015. WIND1-based acquisition of regeneration competency in Arabidopsis and rapeseed. |
| [23] |
Zhao Y, Cheng P, Liu Y, Liu C, Hu Z, et al. 2025. A highly efficient soybean transformation system using GRF3-GIF1 chimeric protein. |
| [24] |
Kong J, Martin-Ortigosa S, Finer J, Orchard N, Gunadi A, et al. 2020. Overexpression of the transcription factor GROWTH-REGULATING FACTOR5 improves transformation of dicot and monocot species. |
| [25] |
Liu X, Bie XM, Lin X, Li M, Wang H, et al. 2023. Uncovering the transcriptional regulatory network involved in boosting wheat regeneration and transformation. |
| [26] |
Wang K, Shi L, Liang X, Zhao P, Wang W, et al. 2022. The gene TaWOX5 overcomes genotype dependency in wheat genetic transformation. |
| [27] |
Liu K, Yang A, Yan J, Liang Z, Yuan G, et al. 2023. MdAIL5 overexpression promotes apple adventitious shoot regeneration by regulating hormone signaling and activating the expression of shoot development-related genes. |
| [28] |
Kang M, Lee K, Finley T, Chappell H, Veena V, et al. 2022. An improved Agrobacterium-mediated transformation and genome-editing method for maize inbred B104 using a ternary vector system and immature embryos. |
| [29] |
Raman V, Rojas CM, Vasudevan B, Dunning K, Kolape J, et al. 2022. Agrobacterium expressing a type III secretion system delivers Pseudomonas effectors into plant cells to enhance transformation. |
| [30] |
Yang F, Li G, Felix G, Albert M, Guo M. 2023. Engineered Agrobacterium improves transformation by mitigating plant immunity detection. |
| [31] |
Demirer GS, Zhang H, Matos JL, Goh NS, Cunningham FJ, et al. 2019. High aspect ratio nanomaterials enable delivery of functional genetic material without DNA integration in mature plants. |
| [32] |
Wang ZP, Zhang ZB, Zheng DY, Zhang TT, Li XL, et al. 2022. Efficient and genotype independent maize transformation using pollen transfected by DNA-coated magnetic nanoparticles. |
| [33] |
Liu Z, Zhang J, Cai Y, Wang H, Luo M, et al. 2024. Improving seed shattering resistance in wild O. alta rice with mesoporous silica nanoparticle delivery systems. |
| [34] |
She L, Cheng X, Jiang P, Shen S, Dai F, et al. 2025. Modified carbon dot-mediated transient transformation for genomic and epigenomic studies in wheat. |
| [35] |
Wu H, Sparks C, Amoah B, Jones HD. 2003. Factors influencing successful Agrobacterium-mediated genetic transformation of wheat. |
| [36] |
Udayabhanu J, Huang T, Xin S, Cheng J, Hua Y, et al. 2022. Optimization of the transformation protocol for increased efficiency of genetic transformation in Hevea brasiliensis. |
| [37] |
Zhang M, Wang Y, Chen X, Xu F, Ding M, et al. 2021. Plasma membrane H+-ATPase overexpression increases rice yield via simultaneous enhancement of nutrient uptake and photosynthesis. |
| [38] |
Wei S, Li X, Lu Z, Zhang H, Ye X, et al. 2022. A transcriptional regulator that boosts grain yields and shortens the growth duration of rice. |
| [39] |
Tao K, Li Y, Hu Y, Li Y, Zhang D, et al. 2023. Overexpression of ZmEXPA5 reduces anthesis-silking interval and increases grain yield under drought and well-watered conditions in maize. |
| [40] |
Wang L, Yang Y, Yang Z, Li W, Hu D, et al. 2023. GmFtsH25 overexpression increases soybean seed yield by enhancing photosynthesis and photosynthates. |
| [41] |
Gao C, Yuan J, Lu J, Ye W, Zhi J, et al. 2025. COL3a simultaneously regulates flowering and branching to improve grain yield in soybean. |
| [42] |
Zhong Y, Wang Y, Pan X, Wang R, Li D, et al. 2025. ZmCCD8 regulates sugar and amino acid accumulation in maize kernels via strigolactone signalling. |
| [43] |
Zhang X, Jia H, Li T, Wu J, Nagarajan R, et al. 2022. TaCol-B5 modifies spike architecture and enhances grain yield in wheat. |
| [44] |
Zhou Y, Chen M, Guo J, Wang Y, Min D, et al. 2020. Overexpression of soybean DREB1 enhances drought stress tolerance of transgenic wheat in the field. |
| [45] |
Maeda S, Yokotani N, Oda K, Mori M. 2020. Enhanced resistance to fungal and bacterial diseases in tomato and Arabidopsis expressing BSR2 from rice. |
| [46] |
Ngaki MN, Sahoo DK, Wang B, Bhattacharyya MK. 2021. Overexpression of a plasma membrane protein generated broad-spectrum immunity in soybean. |
| [47] |
Yuan H, Cheng M, Fan F, Zheng X, Wang R, et al. 2024. OsGRF6-OsYUCCA1/OsWRKY82 signaling cascade upgrade grain yield and bacterial blight resistance in rice. |
| [48] |
Mo H, Chang H, Zhao G, Hu G, Luo X, et al. 2024. iJAZ-based approach to engineer lepidopteran pest resistance in multiple crop species. |
| [49] |
Qiu D, Hu W, Zhou Y, Xiao J, Hu R, et al. 2021. TaASR1-D confers abiotic stress resistance by affecting ROS accumulation and ABA signalling in transgenic wheat. |
| [50] |
Zhang H, Mao L, Xin M, Xing H, Zhang Y, et al. 2022. Overexpression of GhABF3 increases cotton(Gossypium hirsutum L.) tolerance to salt and drought. |
| [51] |
Yu TF, Hou ZH, Wang HL, Chang SY, Song XY, et al. 2024. Soybean steroids improve crop abiotic stress tolerance and increase yield. |
| [52] |
Shang C, Liu X, Chen G, Li G, Hu S, et al. 2024. SlWRKY81 regulates Spd synthesis and Na+/K+ homeostasis through interaction with SlJAZ1 mediated JA pathway to improve tomato saline-alkali resistance. |
| [53] |
Liao M, Ma Z, Kang Y, Zhang B, Gao X, et al. 2023. ENHANCED DISEASE SUSCEPTIBILITY 1 promotes hydrogen peroxide scavenging to enhance rice thermotolerance. |
| [54] |
Mishra D, Shekhar S, Subba P, Keshava Prasad TS, Chakraborty S, et al. 2024. Wheat TaNACα18 functions as a positive regulator of high-temperature adaptive responses and improves cell defense machinery. |
| [55] |
He F, Xu J, Jian Y, Duan S, Hu J, et al. 2023. Overexpression of galactinol synthase 1 from Solanum commersonii (ScGolS1) confers freezing tolerance in transgenic potato. |
| [56] |
Goralogia GS, Willig C, Strauss SH. 2025. Engineering Agrobacterium for improved plant transformation. |
| [57] |
Broothaerts W, Mitchell HJ, Weir B, Kaines S, Smith LMA, et al. 2005. Gene transfer to plants by diverse species of bacteria. |
| [58] |
Dong OX, Ronald PC. 2021. Targeted DNA insertion in plants. |
| [59] |
Vejlupkova Z, Warman C, Sharma R, Scheller HV, Mortimer JC, et al. 2020. No evidence for transient transformation via pollen magnetofection in several monocot species. |
| [60] |
Horstman A, Li M, Heidmann I, Weemen M, Chen B, et al. 2017. The BABY BOOM transcription factor activates the LEC1-ABI3-FUS3-LEC2 network to induce somatic embryogenesis. |
| [61] |
Jha P, Ochatt SJ, Kumar V. 2020. WUSCHEL: a master regulator in plant growth signaling. |
| [62] |
Iwase A, Harashima H, Ikeuchi M, Rymen B, Ohnuma M, et al. 2017. WIND1 promotes shoot regeneration through transcriptional activation of ENHANCER OF SHOOT REGENERATION1 in Arabidopsis. |
| [63] |
Ikeuchi M, Iwase A, Rymen B, Lambolez A, Kojima M, et al. 2017. Wounding triggers callus formation via dynamic hormonal and transcriptional changes. |
| [64] |
Feng Q, Xiao L, He Y, Liu M, Wang J, et al. 2021. Highly efficient, genotype-independent transformation and gene editing in watermelon (Citrullus lanatus) using a chimeric ClGRF4-GIF1 gene. |
| [65] |
Raman V, Mysore KS. 2023. Engineering Agrobacterium tumefaciens with a type III secretion system to express type III effectors. |
| [66] |
Zipfel C, Kunze G, Chinchilla D, Caniard A, Jones JDG, et al. 2006. Perception of the bacterial PAMP EF-Tu by the receptor EFR restricts Agrobacterium-mediated transformation. |
| [67] |
Cunningham FJ, Goh NS, Demirer GS, Matos JL, Landry MP. 2018. Nanoparticle-mediated delivery towards advancing plant genetic engineering. |
| [68] |
Shivashakarappa K, Marriboina S, Dumenyo K, Taheri A, Yadegari Z. 2025. Nanoparticle-mediated gene delivery techniques in plant systems. |
| [69] |
Huynh J, Hotton SK, Chan R, Syed Y, Thomson J. 2022. Evaluation of novel surfactants for plant transformation. |
| [70] |
Hwang HH, Yu M, Lai EM. 2017. Agrobacterium-mediated plant transformation: biology and applications. |
| [71] |
Guan S, Kang X, Ge J, Fei R, Duan S, et al. 2022. An efficient Agrobacterium-mediated transient transformation system and its application in gene function elucidation in Paeonia lactiflora Pall. |
| [72] |
Liu Y, Yang H, Sakanishi A. 2006. Ultrasound: mechanical gene transfer into plant cells by sonoporation. |
| [73] |
Zhang LJ, Cheng LM, Xu N, Zhao NM, Li CG, et al. 1991. Efficient Transformation of Tobacco by Ultrasonication. |
| [74] |
Király A, Farkas D, Dobránszki J. 2025. Ultrasound in plant life and its application perspectives in horticulture and agriculture. |
| [75] |
Wang W, Zhang D, Chu C. 2023. OsDREB1C, an integrator for photosynthesis, nitrogen use efficiency, and early flowering. |
| [76] |
Guan JC, Koch KE, Suzuki M, Wu S, Latshaw S, et al. 2012. Diverse roles of strigolactone signaling in maize architecture and the uncoupling of a branching-specific subnetwork. |
| [77] |
Zhao X, He Y, Liu Y, Wang Z, Zhao J. 2024. JAZ proteins: key regulators of plant growth and stress response. |
| [78] |
Guo W, Zhang J, Zhang N, Xin M, Peng H, et al. 2015. The wheat NAC transcription factor TaNAC2L is regulated at the transcriptional and post-translational levels and promotes heat stress tolerance in transgenic Arabidopsis. |
| [79] |
Abdul Aziz M, Brini F, Rouached H, Masmoudi K. 2022. Genetically engineered crops for sustainably enhanced food production systems. |
| [80] |
Bekele-Alemu A, Dessalegn-Hora O, Safawo-Jarso T, Ligaba-Osena A. 2025. Rethinking progress: harmonizing the discourse on genetically modified crops. |
| [81] |
Cheng X, Li H, Tang Q, Zhang H, Liu T, et al. 2024. Trends in the global commercialization of genetically modified crops in 2023. |
| [82] |
MacDonald JM, Dong X, Fuglie K. 2023. Concentration and competition in U.S. agribusiness. Washington, DC, United States: Department of Agriculture, Economic Research Service. 57 pp. doi: 10.32747/2023.8054022.ers |
| [83] |
Sun Z, Scherer L, Tukker A, Behrens P. 2020. Linking global crop and livestock consumption to local production hotspots. |
| [84] |
Hu N, Tian H, Li Y, Li X, Li D, et al. 2025. pHNRhCas9NG, single expression cassette-based dual-component dual-transcription unit CRISPR/Cas9 system for plant genome editing. |
| [85] |
He Y, Han Y, Ma Y, Liu S, Fan T, et al. 2024. Expanding plant genome editing scope and profiles with CRISPR-FrCas9 systems targeting palindromic TA sites. |
| [86] |
Li B, Shang Y, Wang L, Lv J, Wu Q, et al. 2025. Efficient genome editing in dicot plants using calreticulin promoter-driven CRISPR/Cas system. |
| [87] |
Liu Q, Zhao C, Sun K, Deng Y, Li Z. 2023. Engineered biocontainable RNA virus vectors for non-transgenic genome editing across crop species and genotypes. |
| [88] |
Qiao JH, Zang Y, Gao Q, Liu S, Zhang XW, et al. 2025. Transgene- and tissue culture-free heritable genome editing using RNA virus-based delivery in wheat. |
| [89] |
Nagy B, Öktem A, Ferenc G, Ungor D, Kalac A, et al. 2023. CRISPR/Cas9 mutagenesis through introducing a nanoparticle complex made of a cationic polymer and nucleic acids into maize protoplasts. |