[1]

Zhang W, Yu J, Xu Y, Wang Z, Liu L, et al. 2021. Alternate wetting and drying irrigation combined with the proportion of polymer-coated urea and conventional urea rates increases grain yield, water and nitrogen use efficiencies in rice. Field Crops Research 268:108165

doi: 10.1016/j.fcr.2021.108165
[2]

National Bureau of Statistics of China. 2022. China Statistical Yearbook. www.stats.gov.cn/gk

[3]

Lai Z, Fan J, Yang R, Xu X, Liu L, et al. 2022. Interactive effects of plant density and nitrogen rate on grain yield, economic benefit, water productivity and nitrogen use efficiency of drip-fertigated maize in northwest China. Agricultural Water Management 263:107453

doi: 10.1016/j.agwat.2021.107453
[4]

Zhao L, Tang Q, Song Z, Yin Y, Wang G, et al. 2023. Increasing the yield of drip-irrigated rice by improving photosynthetic performance and enhancing nitrogen metabolism through optimizing water and nitrogen management. Frontiers in Plant Science 14:1075625

doi: 10.3389/fpls.2023.1075625
[5]

Pandey V, Shukla A. 2015. Acclimation and tolerance strategies of rice under drought stress. Rice Science 22:147−161

doi: 10.1016/j.rsci.2015.04.001
[6]

Tuong P, Bouman BAM, Mortimer M. 2005. More rice, less water—integrated approaches for increasing water productivity in irrigated rice-based systems in Asia. Plant Production Science 8:231−241

doi: 10.1626/pps.8.231
[7]

Bai R, Chen L, Zhang X, Wei G, Wei C. 2017. Effect of salinity and soil temperature on the growth and physiology of drip-irrigated rice seedlings. Archives of Agronomy and Soil Science 63:513−524

doi: 10.1080/03650340.2016.1223842
[8]

Zhang X, Liu H, Meng C, Zhang Z, Wang M, et al. 2019. Ammonium alleviates iron deficiency of drip-irrigated rice seedlings in low soil temperature in calcareous soil. Acta Agriculturae Scandinavica, Section B — Soil & Plant Science 69:411−421

doi: 10.1080/09064710.2019.1590626
[9]

Tian Y, Shi X, Shi F, Zhang H, Liang Q, et al. 2023. Late nitrogen fertilization increases biomass of cotton bolls by reinforcing source—sink performance. Industrial Crops and Products 206:117663

doi: 10.1016/j.indcrop.2023.117663
[10]

Lyu W, Su JW, Yu XY, Yue NY, Dong ZC, et al. 2023. Physiological mechanism of the effects of two nitrogen forms on yield and photosynthetic characteristics of rice. Journal of Southern Agriculture 54:1780−1788 (in Chinese)

doi: 10.3969/j.issn.2095-1191.2023.06.019
[11]

Duan YH, Zhang YL, Shen QR. 2004. Nitrification in rice rhizosphere and the nitrate nutrition of rice. Acta Pedologica Sinica 41:803−809 (in Chinese)

doi: 10.3321/j.issn:0564-3929.2004.05.021
[12]

Kirk GJD. 2001. Plant-mediated processess to acquire nutrients: nitrogen uptake by rice plants. Plant and Soil 232:129−134

doi: 10.1023/A:1010341116376
[13]

Xu GW, Jiang MM, Lu DK, Wang HZ, Chen MC. 2020. Nitrogen forms affect the root characteristic, photosynthesis, grain yield, and nitrogen use efficiency of rice under different irrigation regimes. Crop Science 60:2594−2610

doi: 10.1002/csc2.20242
[14]

Zheng E, Yang H, Zhang Z. 2018. Influence of different nitrogen forms application on rice photosynthesis: fluorescence with water-saving irrigation in black soil region of Songnen Plain, Northeast China. Paddy and Water Environment 16:795−804

doi: 10.1007/s10333-018-0670-y
[15]

Li K, Li H, Shi G, Xiao M, Li C, et al. 2020. Physiological responses of Scirpus validus to nitrate stress. Polish Journal of Environmental Studies 29:163−172

doi: 10.15244/pjoes/99903
[16]

Xie Y, Lv Y, Jia L, Zheng L, Li Y, et al. 2023. Plastid-localized amino acid metabolism coordinates rice ammonium tolerance and nitrogen use efficiency. Nature Plants 9:1514−1529

doi: 10.1038/s41477-023-01494-x
[17]

de Souza EACC, Alvarez-Pizarro JC, de Sousa Lopes L, de Souza Miranda R, Gomes-Filho E. 2021. Nitrate and ammonium nutrition modulates the photosynthetic performance and antioxidant defense in salt-stressed grass species. Journal of Soil Science and Plant Nutrition 21:3016−3029

doi: 10.1007/s42729-021-00586-x
[18]

Zeng J, Tang J, Zhang F, Wang Y, Kang H, et al. 2021. Ammonium regulates redox homeostasis and photosynthetic ability to mitigate copper toxicity in wheat seedlings. Ecotoxicology and Environmental Safety 226:112825

doi: 10.1016/j.ecoenv.2021.112825
[19]

Heuermann D, Hahn H, von Wirén N. 2021. Seed yield and nitrogen efficiency in oilseed rape after ammonium nitrate or urea fertilization. Frontiers in Plant Science 11:608785

doi: 10.3389/fpls.2020.608785
[20]

Krouk G. 2016. Hormones and nitrate: a two-way connection. Plant Molecular Biology 91:599−606

doi: 10.1007/s11103-016-0463-x
[21]

Tang Q, Wang G, Zhao L, Song Z, Li Y. 2025. Responses of yield, root traits and their plasticity to the nitrogen environment in nitrogen-efficient cultivars of drip-irrigated rice. Journal of Integrative Agriculture 24:480−496

doi: 10.1016/j.jia.2023.12.014
[22]

van Kooten O, Snel JFH. 1990. The use of chlorophyll fluorescence nomenclature in plant stress physiology. Photosynthesis Research 25:147−150

doi: 10.1007/BF00033156
[23]

Qiu ZB, Li JT, Yue M. 2010. The damage repair role of He-Ne laser on wheat exposed to osmotic stress. Canadian Journal of Plant Science 90:691−698

doi: 10.4141/cjps09118
[24]

Kant S, Seneweera S, Rodin J, Materne M, Burch D, et al. 2012. Improving yield potential in crops under elevated CO2: integrating the photosynthetic and nitrogen utilization efficiencies. Frontiers in Plant Science 3:162

doi: 10.3389/fpls.2012.00162
[25]

Tanaka R, Tanaka A. 2011. Chlorophyll cycle regulates the construction and destruction of the light-harvesting complexes. Biochimica et Biophysica Acta (BBA) - Bioenergetics 1807:968−976

doi: 10.1016/j.bbabio.2011.01.002
[26]

Li Y, Gu X, Li Y, Fang H, Chen P. 2023. Ridge-furrow mulching combined with appropriate nitrogen rate for enhancing photosynthetic efficiency, yield and water use efficiency of summer maize in a semi-arid region of China. Agricultural Water Management 287:108450

doi: 10.1016/j.agwat.2023.108450
[27]

Kasajima I. 2017. Difference in oxidative stress tolerance between rice cultivars estimated with chlorophyll fluorescence analysis. BMC Research Notes 10:168

doi: 10.1186/s13104-017-2489-9
[28]

Song N, Guo SW, Shen QR. 2007. Effects of different nitrogen forms and water stress on water absorption, photosynthesis and growth of Oryza sativa Seedlings. Chinese Bulletin of Botany 24:477−483 (in Chinese)

doi: 10.3969/j.issn.1674-3466.2007.04.007
[29]

Li CD, Dong HR, Li JC. 2003. Influence of various ratios of nitrogen nutrition on photosynthetic and sugar metabolism of cotton. Cotton Science 15:87−90 (in Chinese)

doi: 10.3969/j.issn.1002-7807.2003.02.005
[30]

Guo S, Chen G, Zhou Y, Shen Q. 2007. Ammonium nutrition increases photosynthesis rate under water stress at early development stage of rice (Oryza sativa L.). Plant and Soil 296:115−124

doi: 10.1007/s11104-007-9302-9
[31]

Guo S, Zhou Y, Shen Q, Zhang F. 2007. Effect of ammonium and nitrate nutrition on some physiological processes in higher plants - growth, photosynthesis, photorespiration, and water relations. Plant Biology 9:21−29

doi: 10.1055/s-2006-924541
[32]

Guo HX, Liu WQ, Shi YC. 2006. Effects of different nitrogen forms on photosynthetic rate and the chlorophyll fluorescence induction kinetics of flue-cured tobacco. Photosynthetica 44:140−142

doi: 10.1007/s11099-005-0170-3
[33]

Zhou YH, Zhang YL, Wang XM, Cui JX, Xia XJ, et al. 2011. Effects of nitrogen form on growth, CO2 assimilation, chlorophyll fluorescence, and photosynthetic electron allocation in cucumber and rice plants. Journal of Zhejiang University SCIENCE B 12:126−134

doi: 10.1631/jzus.B1000059
[34]

Wei HY, Zhang HC, Ma Q, Dai QG, Huo ZY, et al. 2009. Photosynthetic characteristics of flag leaf in rice genotypes with different nitrogen use efficiencies. Acta Agronomica Sinica 35:2243−2251

doi: 10.3724/sp.j.1006.2009.02243
[35]

Lee S, Masclaux-Daubresse C. 2021. Current understanding of leaf senescence in rice. International Journal of Molecular Sciences 22:4515

doi: 10.3390/ijms22094515
[36]

Singh M, Singh VP, Prasad SM. 2016. Nitrogen modifies NaCl toxicity in eggplant seedlings: assessment of chlorophyll a fluorescence, antioxidative response and proline metabolism. Biocatalysis and Agricultural Biotechnology 7:76−86

doi: 10.1016/j.bcab.2016.05.007
[37]

Walch-Liu P, Neumann G, Bangerth F, Engels C. 2000. Rapid effects of nitrogen form on leaf morphogenesis in tobacco. Journal of Experimental Botany 51:227−237

doi: 10.1093/jexbot/51.343.227
[38]

Chen H, Zhang Q, Cai H, Zhou W, Xu F. 2018. H2O2 mediates nitrate-induced iron chlorosis by regulating iron homeostasis in rice. Plant, Cell & Environment 41:767−781

doi: 10.1111/pce.13145
[39]

Hejazi Z, Safi S, Barakzai AL, Karimi BK, Quraishi FM, et al. 2023. Cultivar effect on root development in pomegranate hardwood cuttings treated with auxin. Rhizosphere 25:100661

doi: 10.1016/j.rhisph.2022.100661
[40]

Wu H, Xiang J, Zhang Y, Zhang Y, Peng S, et al. 2018. Effects of post-anthesis nitrogen uptake and translocation on photosynthetic production and rice yield. Scientific Reports 8:12891

doi: 10.1038/s41598-018-31267-y
[41]

Kim M, Lee B, Sung J. 2025. CN metabolism and nitrogen use efficiency of rice with different nitrogen form and rate. PLoS One 20:e0318522

doi: 10.1371/journal.pone.0318522
[42]

O'Brien JA, Vega A, Bouguyon E, Krouk G, Gojon A, et al. 2016. Nitrate transport, sensing, and responses in plants. Molecular Plant 9:837−856

doi: 10.1016/j.molp.2016.05.004
[43]

Subbarao GV, Searchinger TD. 2021. Opinion: a “more ammonium solution” to mitigate nitrogen pollution and boost crop yields. Proceedings of the National Academy of Sciences of the United States of America 118:e2107576118

doi: 10.1073/pnas.2107576118
[44]

Fu H, Cui D, Shen H. 2021. Effects of nitrogen forms and application rates on nitrogen uptake, photosynthetic characteristics and yield of double-cropping rice in south China. Agronomy 11:158

doi: 10.3390/agronomy11010158
[45]

Wang ZH, Miao YF, Li SX. 2015. Effect of ammonium and nitrate nitrogen fertilizers on wheat yield in relation to accumulated nitrate at different depths of soil in drylands of China. Field Crops Research 183:211−224

doi: 10.1016/j.fcr.2015.07.019
[46]

Tegeder M, Masclaux-Daubresse C. 2018. Source and sink mechanisms of nitrogen transport and use. New Phytologist 217:35−53

doi: 10.1111/nph.14876
[47]

Yan Y, Zhang Z, Sun H, Liu X, Xie J, et al. 2023. Nitrate confers rice adaptation to high ammonium by suppressing its uptake but promoting its assimilation. Molecular Plant 16:1871−1874

doi: 10.1016/j.molp.2023.11.008
[48]

Meng X, Li Y, Yao H, Wang J, Dai F, et al. 2020. Nitrification and urease inhibitors improve rice nitrogen uptake and prevent denitrification in alkaline paddy soil. Applied Soil Ecology 154:103665

doi: 10.1016/j.apsoil.2020.103665
[49]

Duan M, Han C, Zhang X, Wei Z, Wang Z, et al. 2025. Spatial and temporal dynamics of photosynthetically active radiation in crops: effects of canopy structure on yield. Agronomy 15:940

doi: 10.3390/agronomy15040940
[50]

Zhang G, Li Z, Zhu Q, Yang C, Shu H, et al. 2025. Cropping patterns and plant population density alter nitrogen partitioning among photosynthetic components, leaf photosynthetic capacity and photosynthetic nitrogen use efficiency in field-grown soybean. Industrial Crops and Products 226:120680

doi: 10.1016/j.indcrop.2025.120680