[1]

Mostafa S, Wang Y, Zeng W, Jin B. 2022. Floral scents and fruit aromas: functions, compositions, biosynthesis, and regulation. Frontiers in Plant Science 13:860157

doi: 10.3389/fpls.2022.860157
[2]

Schiestl FP. 2010. The evolution of floral scent and insect chemical communication. Ecology Letters 13:643−656

doi: 10.1111/j.1461-0248.2010.01451.x
[3]

Gross K, Sun M, Schiestl FP. 2016. Why do floral perfumes become different? region-specific selection on floral scent in a terrestrial orchid. PLoS One 11:e0147975

doi: 10.1371/journal.pone.0147975
[4]

Pichersky E, Noel JP, Dudareva N. 2006. Biosynthesis of plant volatiles: nature's diversity and ingenuity. Science 311:808−811

doi: 10.1126/science.1118510
[5]

Knudsen JT, Eriksson R, Gershenzon J, Ståhl B. 2006. Diversity and distribution of floral scent. The Botanical Review 72:1

doi: 10.1663/0006-8101(2006)72[1:DADOFS]2.0.CO;2
[6]

Abbas F, Zhou Y, O'Neill Rothenberg D, Alam I, Ke Y, et al. 2023. Aroma components in horticultural crops: chemical diversity and usage of metabolic engineering for industrial applications. Plants 12(9):1748

doi: 10.3390/plants12091748
[7]

Farré-Armengol G, Filella I, Llusià J, Peñuelas J. 2017. β-ocimene, a key floral and foliar volatile involved in multiple interactions between plants and other organisms. Molecules 22(7):1148

doi: 10.3390/molecules22071148
[8]

Fujihashi M, Sato T, Tanaka Y, Yamamoto D, Nishi T, et al. 2018. Crystal structure and functional analysis of large-terpene synthases belonging to a newly found subclass. Chemical Science 9:3754−3758

doi: 10.1039/C8SC00289D
[9]

Huang Y, Xie FJ, Cao X, Li MY. 2021. Research progress in biosynthesis and regulation of plant terpenoids. Biotechnology & Biotechnological Equipment 35:1799−1808

doi: 10.1080/13102818.2021.2020162
[10]

Lange BM, Croteau R. 1999. Isoprenoid biosynthesis via a mevalonate-independent pathway in plants: cloning and heterologous expression of 1-deoxy-D-xylulose-5-phosphate reductoisomerase from peppermint. Archives of Biochemistry and Biophysics 365:170−174

doi: 10.1006/abbi.1999.1168
[11]

Eisenreich W, Bacher A, Arigoni D, Rohdich F. 2004. Biosynthesis of isoprenoids via the non-mevalonate pathway. Cellular and Molecular Life Sciences CMLS 61:1401−1426

doi: 10.1007/s00018-004-3381-z
[12]

Chaurasiya ND, Sangwan NS, Sabir F, Misra L, Sangwan RS. 2012. Withanolide biosynthesis recruits both mevalonate and DOXP pathways of isoprenogenesis in Ashwagandha Withania somnifera L. (Dunal). Plant Cell Reports 31:1889−1897

doi: 10.1007/s00299-012-1302-4
[13]

Yadav RK, Sangwan RS, Sabir F, Srivastava AK, Sangwan NS. 2014. Effect of prolonged water stress on specialized secondary metabolites, peltate glandular trichomes, and pathway gene expression in Artemisia annua L. Plant Physiology and Biochemistry 74:70−83

doi: 10.1016/j.plaphy.2013.10.023
[14]

Bick JA, Lange BM. 2003. Metabolic cross talk between cytosolic and plastidial pathways of isoprenoid biosynthesis: unidirectional transport of intermediates across the chloroplast envelope membrane. Archives of Biochemistry and Biophysics 415:146−154

doi: 10.1016/S0003-9861(03)00233-9
[15]

Flügge UI, Gao W. 2005. Transport of isoprenoid intermediates across chloroplast envelope membranes. Plant Biology 7:91−97

doi: 10.1055/s-2004-830446
[16]

Ward JL, Baker JM, Llewellyn AM, Hawkins ND, Beale MH. 2011. Metabolomic analysis of Arabidopsis reveals hemiterpenoid glycosides as products of a nitrate ion-regulated, carbon flux overflow. Proceedings of the National Academy of Sciences of the United States of America 108:10762−10767

doi: 10.3410/f.11426956.12894057
[17]

Guirimand G, Guihur A, Phillips MA, Oudin A, Glévarec G, et al. 2012. A single gene encodes isopentenyl diphosphate isomerase isoforms targeted to plastids, mitochondria and peroxisomes in Catharanthus roseus. Plant Molecular Biology 79:443−459

doi: 10.1007/s11103-012-9923-0
[18]

Chappell J. 2002. The genetics and molecular genetics of terpene and sterol origami. Current Opinion in Plant Biology 5:151−157

doi: 10.1016/S1369-5266(02)00241-8
[19]

Qin W, Li Y, Liu H, Yan X, Hu X, et al. 2024. Integrated multi-omics profiling reveals a landscape of dramatic metabolic defect in Artemisia annua. Horticulture Research 11:uhae174

doi: 10.1093/hr/uhae174
[20]

Yeon JY, Kim WS. 2021. Biosynthetic linkage between the color and scent of flowers: a review. Horticultural Science and Technology 39:697−713

doi: 10.7235/HORT.20210062
[21]

Feng Z, Li Y, Li M, Wang Y, Zhang L, et al. 2019. Tea aroma formation from six model manufacturing processes. Food Chemistry 285:347−354

doi: 10.1016/j.foodchem.2019.01.174
[22]

Yue R, Li Y, Qi Y, Liang X, Zheng Z, et al. 2025. Divergent MYB paralogs determine spatial distribution of linalool mediated by JA and DNA demethylation participating in aroma formation and cold tolerance of tea plants. Plant Biotechnology Journal 23:1455−1475

doi: 10.1111/pbi.14598
[23]

Chen Y, Lu X, Gao T, Zhou Y. 2025. The scent of lily flowers: advances in the identification, biosynthesis, and regulation of fragrance components. International Journal of Molecular Sciences 26(2):468

doi: 10.3390/ijms26020468
[24]

Hu Z, Tang B, Wu Q, Zheng J, Leng P, et al. 2017. Transcriptome sequencing analysis reveals a difference in monoterpene biosynthesis between scented Lilium 'Siberia' and unscented Lilium 'Novano'. Frontiers in Plant Science 8:1351

doi: 10.3389/fpls.2017.01351
[25]

Kong Y, Bai J, Lang L, Bao F, Dou X, et al. 2017. Variation in floral scent compositions of different lily hybrid groups. Journal of the American Society for Horticultural Science 142:175−183

doi: 10.21273/JASHS03934-16
[26]

Wang T, Fan J, Zhang H, Fan W, Du F. 2018. Correlation of monoterpenes release and LhTPS expression in Lilium plants. Journal of Henan Agricultral Sciences 47:88−108
[27]

Cao L, Jiang F, Liu D, Zhang J, Yang T, et al. 2025. Genome-wide characterization of differentially expressed scent genes in the MEP control network of the flower of Lilium 'Sorbonne'. Molecular Biotechnology 67:510−526

doi: 10.1007/s12033-024-01063-3
[28]

Yu J, Wu N, Fu R, Xue L, Zhang J, et al. 2024. Cloning and expression characterisation of LiDXS2 gene in Oriental Lily (Lilium 'Sorbonne'). Folia Horticulturae 36:463−474

doi: 10.2478/fhort-2024-0030
[29]

Liu C, Zhou X, Li Y, Hittinger CT, Pan R, et al. 2024. The influence of the number of tree searches on maximum likelihood inference in phylogenomics. Systematic Biology 73:807−822

doi: 10.1093/sysbio/syae031
[30]

Álvarez-Urdiola R, Bustamante M, Ribes J, Riechmann JL. 2023. Gene expression analysis by quantitative real-time PCR for floral tissues. In Flower Development, eds. Riechmann JL, Ferrándiz C. New York, NY: Humana. pp. 403−428 doi: 10.1007/978-1-0716-3299-4_20
[31]

Carlson ED, Rajniak J, Sattely ES. 2023. Multiplicity of the Agrobacterium infection of Nicotiana benthamiana for transient DNA delivery. ACS Synthetic Biology 12:2329−2338

doi: 10.1021/acssynbio.3c00148
[32]

Sun G, Ju Y, Zhang C, Li L, Jiang X, et al. 2024. Styrax japonicus functional genomics: an efficient virus induced gene silencing (VIGS) system. Horticultural Plant Journal 10:252−258

doi: 10.1016/j.hpj.2023.05.003
[33]

Tong Z, Dong T, Li Q, Wang R, He J, et al. 2024. R2R3-type LoMYB21 affects jasmonate-regulated development and dehiscence of anthers in lily (Lilium oriental hybrids). Horticultural Plant Journal 10:239−251

doi: 10.1016/j.hpj.2023.04.004
[34]

Li M, Yang Y, Raza A, Yin S, Wang H, et al. 2021. Heterologous expression of Arabidopsis thaliana rty gene in strawberry (Fragaria × ananassa Duch.) improves drought tolerance. BMC Plant Biology 21:57

doi: 10.21203/rs.3.rs-34565/v2
[35]

Mase K, Kamiya Y, Sakaoka S, Morikami A, Tsukagoshi H. 2025. Root growth control by negative regulation of MYB50 under ABA signaling in Arabidopsis. Bioscience, Biotechnology, and Biochemistry 89:362−370

doi: 10.1093/bbb/zbae195
[36]

Wang F, Liang Z, Ma X, He Z, Li J, et al. 2024. LcMPK3 and LcMPK6 positively regulate fruitlet abscission in litchi. Molecular Horticulture 4:29

doi: 10.1186/s43897-024-00109-z
[37]

Ershova NM, Sheshukova EV, Kamarova KA, Alimova AR, Savchenko YY, et al. 2025. Xyloglucan endotransglycosylase/hydrolase downregulation increases Nicotiana benthamiana tolerance to tobacco mosaic virus infection. International Journal of Molecular Sciences 26(22):11183

doi: 10.3390/ijms262211183
[38]

Ilbeigi V, Valadbeigi Y, Slováková L, Matejčík Š. 2022. Solid Phase microextraction–multicapillary column–ion mobility spectrometry (SPME–MCC–IMS) for detection of methyl salicylate in tomato leaves. Journal of Agricultural and Food Chemistry 70:15593−15601

doi: 10.1021/acs.jafc.2c05570
[39]

Rontani JF. 2022. Use of gas chromatography-mass spectrometry techniques (GC-MS, GC-MS/MS and GC-QTOF) for the characterization of photooxidation and autoxidation products of lipids of autotrophic organisms in environmental samples. Molecules 27(5):1629

doi: 10.3390/molecules27051629
[40]

Zhou Y, Wang F, Wang Q, Chen Y, Dong J, et al. 2024. Diversity of chemical compounds in lily fragrance and identification of key sensory markers. Industrial Crops and Products 222:120034

doi: 10.1016/j.indcrop.2024.120034
[41]

Liu Z, Wang Y, Xu H, Zhu J, Zhai H, et al. 2025. Integrative analyses of metabolome and transcriptome reveal the regulatory network underlying flavor differences in lychee pulp from different varieties. Plant Physiology and Biochemistry 229:110594

doi: 10.1016/j.plaphy.2025.110594
[42]

Lipko A, Pączkowski C, Perez-Fons L, Fraser PD, Kania M, et al. 2023. Divergent contribution of the MVA and MEP pathways to the formation of polyprenols and dolichols in Arabidopsis. Biochemical Journal 480:495−520

doi: 10.1042/BCJ20220578
[43]

Zhang T, Guo Y, Shi X, Yang Y, Chen J, et al. 2020. Overexpression of LiTPS2 from a cultivar of lily (Lilium 'Siberia') enhances the monoterpenoids content in tobacco flowers. Plant Physiology and Biochemistry 151:391−399

doi: 10.1016/j.plaphy.2020.03.048
[44]

Song C, Ma H, Li R, Zhao G, Niu T, et al. 2024. Analysis of the emitted pattern of floral volatiles and cloning and functional analysis of the PsuLIS gene in tree peony cultivar 'High Noon'. Scientia Horticulturae 326:112750

doi: 10.1016/j.scienta.2023.112750
[45]

Xu Y, Zhang H, Wu Y, Xu W, Cai M, et al. 2024. Functional characterization of LcTPS1 and LcTPS14 explains the biosynthesis of citronellol, citronellal and linalool in Lagerstroemia caudata. Industrial Crops and Products 209:118033

doi: 10.1016/j.indcrop.2024.118033
[46]

Yang Y, Liu X, Hu Z, Zhang Y, Yu H, et al. 2024. Negative regulatory role of LiSRM1 in monoterpene biosynthesis in Lilium 'Siberia'. Plant Physiology and Biochemistry 216:109167

doi: 10.1016/j.plaphy.2024.109167
[47]

Hsieh MH, Chang CY, Hsu SJ, Chen JJ. 2008. Chloroplast localization of methylerythritol 4-phosphate pathway enzymes and regulation of mitochondrial genes in ispD and ispE albino mutants in Arabidopsis. Plant Molecular Biology 66:663−673

doi: 10.1007/s11103-008-9297-5
[48]

Yang X, Chen H, Chen C, Gu G, Zhang B, et al. 2025. Analysis of mechanisms underlying accelerated plant growth induced by NtGLK85 overexpression in tobacco. Scientific Reports 15:40584

doi: 10.1038/s41598-025-24323-x
[49]

Jang S, Li HY. 2018. Overexpression of OsAP2 and OsWRKY24 in Arabidopsis results in reduction of plant size. Plant Biotechnology 35:273−279

doi: 10.5511/plantbiotechnology.18.0508a
[50]

Fan B, Cong D, Sheng H, Zhao K, Zhang T, et al. 2019. Phenotype and potassium uptake of transgenic ZmHAK1 Arabidopsis thaliana. Journal of Jilin Agricultural University 41:149−153 (in Chinese)

doi: 10.13327/j.jjlau.2019.4897
[51]

Jiang F, Liu D, Dai J, Yang T, Zhang J, et al. 2024. Cloning and functional characterization of 2-C-methyl-D-erythritol-4-phosphate cytidylyltransferase (LiMCT) gene in oriental lily (Lilium 'Sorbonne'). Molecular Biotechnology 66:56−67

doi: 10.1007/s12033-023-00729-8
[52]

Estévez JM, Cantero A, Reindl A, Reichler S, León P. 2001. 1-deoxy-D-xylulose-5-phosphate synthase, a limiting enzyme for plastidic isoprenoid biosynthesis in plants. Journal of Biological Chemistry 276:22901−22909

doi: 10.1074/jbc.M100854200
[53]

Khozaei M, Fisk S, Lawson T, Gibon Y, Sulpice R, et al. 2015. Overexpression of plastid transketolase in tobacco results in a thiamine auxotrophic phenotype. The Plant Cell 27:432−447

doi: 10.1105/tpc.114.131011
[54]

Cao J, Jin Q, Kuang J, Wang Y, Xu Y. 2021. Regulation of flowering timing by ABA-NnSnRK1 signaling pathway in lotus. International Journal of Molecular Sciences 22(8):3932

doi: 10.3390/ijms22083932
[55]

Yaish MW, El-Kereamy A, Zhu T, Beatty PH, Good AG, et al. 2010. The APETALA-2-like transcription factor OsAP2-39 controls key interactions between abscisic acid and gibberellin in rice. PLoS Genetics 6:e1001098

doi: 10.1371/journal.pgen.1001098
[56]

Cermanová K, Bublavá P, Darbandsari M, Fellner M, Novák O, et al. 2025. Metabolic network divergence: polyamine and ethylene biosynthesis dynamics in Arabidopsis thaliana and Solanum lycopersicum. Plant Stress 18:101124

doi: 10.1016/j.stress.2025.101124
[57]

Bénédetti H, Vallée B. 2023. LIM kinases: from molecular to pathological features. Cells 12(12):1649

doi: 10.3390/cells12121649
[58]

Hu Q, Zhang Y, Tu Z, Wen S, Wang J, et al. 2024. The identification and functional characterization of the LcMCT gene from Liriodendron chinense reveals its potenatial role in carotenoids biosyanthesis. Gene 902:148180

doi: 10.1016/j.gene.2024.148180