[1]

Bi JX, Wu MJ, Zhang Y, Zhang CM. 2023. Analysis of ascorbic acid in jujube/sour jujube and identification of key function genes. Journal of Fruit Science 40:1823−1831 (in Chinese)

doi: 10.13925/j.cnki.gsxb.20230131
[2]

Liu X, Wu R, Bulley SM, Zhong C, Li D. 2022. Kiwifruit MYBS1-like and GBF3 transcription factors influence L-ascorbic acid biosynthesis by activating transcription of GDP-L-galactose phosphorylase 3. New Phytologist 234:1782−1800

doi: 10.1111/nph.18097
[3]

Liu MJ, Zhao J, Cai QL, Liu GC, Wang JR, et al. 2014. The complex jujube genome provides insights into fruit tree biology. Nature Communications 5:5315

doi: 10.1038/ncomms6315
[4]

Zhang C, Huang J, Li X. 2016. Transcriptomic analysis reveals the metabolic mechanism of L-ascorbic acid in Ziziphus jujuba Mill. Frontiers in Plant Science 7:122

doi: 10.3389/fpls.2016.00122
[5]

Imai T, Ban Y, Terakami S, Yamamoto T, Moriguchi T. 2009. L-Ascorbate biosynthesis in peach: cloning of six L-galactose pathway-related genes and their expression during peach fruit development. Physiologia Plantarum 136:139−149

doi: 10.1111/j.1399-3054.2009.01213.x
[6]

Li M, Ma F, Liang D, Li J, Wang Y. 2010. Ascorbate biosynthesis during early fruit development is the main reason for its accumulation in kiwi. PLoS One 5:e14281

doi: 10.1371/journal.pone.0014281
[7]

Lei D, Lin Y, Chen Q, Zhao B, Tang H, et al. 2022. Transcriptomic analysis and the effect of maturity stage on fruit quality reveal the importance of the L-galactose pathway in the ascorbate biosynthesis of hardy kiwifruit (Actinidia arguta). International Journal of Molecular Sciences 23:6816

doi: 10.3390/ijms23126816
[8]

Wang Y, Diao S, Li H, Ye L, Suo Y, et al. 2023. Comparative metabolomic and transcriptomic analyses reveal distinct ascorbic acid (AsA) accumulation patterns between PCA and PCNA persimmon developing fruit. International Journal of Molecular Sciences 24:15362

doi: 10.3390/ijms242015362
[9]

Wheeler GL, Jones MA, Smirnoff N. 1998. The biosynthetic pathway of vitamin C in higher plants. Nature 393:365−369

doi: 10.1038/30728
[10]

Wolucka BA, Van Montagu M. 2003. GDP-mannose 3', 5'-epimerase forms GDP-L-gulose, a putative intermediate for the de novo biosynthesis of vitamin C in plants. The Journal of Biological Chemistry 278:47483−47490

doi: 10.1074/jbc.M309135200
[11]

Alós E, Rey F, Gil JV, Rodrigo MJ, Zacarias L. 2021. Ascorbic acid content and transcriptional profiling of genes involved in its metabolism during development of petals, leaves, and fruits of orange (Citrus sinensis cv. Valencia Late). Plants 10:2590

doi: 10.3390/plants10122590
[12]

Zhang C, Liu J, Zhang Y, Cai X, Gong P, et al. 2011. Overexpression of SlGMEs leads to ascorbate accumulation with enhanced oxidative stress, cold, and salt tolerance in tomato. Plant Cell Reports 30:389−98

doi: 10.1007/s00299-010-0939-0
[13]

Mounet-Gilbert L, Dumont M, Ferrand C, Bournonville C, Monier A, et al. 2016. Two tomato GDP-D-mannose epimerase isoforms involved in ascorbate biosynthesis play specific roles in cell wall biosynthesis and development. Journal of Experimental Botany 67:4767−4777

doi: 10.1093/jxb/erw260
[14]

Su L, Zhang T, Max Cheng ZM. 2025. Identification of transcription factors contributing to vitamin C synthesis during Rosa roxburghii fruit development by integrating transcriptomics and metabolomics. Horticultural Plant Journal 11:123−132

doi: 10.1016/j.hpj.2023.05.023
[15]

Liu X, Li Y, Zhang X, Xie X, Muzahid ANM, et al. 2025. AcABI5a integrates abscisic acid signaling to developmentally modulate fruit ascorbic acid biosynthesis in kiwifruit. Horticulture Research 12:uhaf111

doi: 10.1093/hr/uhaf111
[16]

Hu T, Ye J, Tao P, Li H, Zhang J, et al. 2016. The tomato HD-Zip I transcription factor SlHZ24 modulates ascorbate accumulation through positive regulation of the ᴅ-mannose/ʟ-galactose pathway. The Plant Journal 85:16−29

doi: 10.1111/tpj.13085
[17]

Yu TF, Hou ZH, Wang HL, Chang SY, Song XY, et al. 2024. Soybean steroids improve crop abiotic stress tolerance and increase yield. Plant Biotechnology Journal 22:2333−2347

doi: 10.1111/pbi.14349
[18]

Wang J, Li G, Li C, Zhang C, Cui L, et al. 2021. NF-Y plays essential roles in flavonoid biosynthesis by modulating histone modifications in tomato. New Phytologist 229:3237−3252

doi: 10.1111/nph.17112
[19]

Chen W, Hu T, Ye J, Wang B, Liu G, et al. 2020. A CCAAT-binding factor, SlNFYA10, negatively regulates ascorbate accumulation by modulating the ᴅ-mannose/ʟ-galactose pathway in tomato. Horticulture Research 7:200

doi: 10.1038/s41438-020-00418-6
[20]

Tao J, Wu H, Li Z, Huang C, Xu X. 2018. Molecular evolution of GDP-D-mannose epimerase (GME), a key gene in plant ascorbic acid biosynthesis. Frontiers in Plant Science 9:1293

doi: 10.3389/fpls.2018.01293
[21]

Zhang C, Geng Y, Liu H, Wu M, Bi J, et al. 2023. Low-acidity ALUMINUM-DEPENDENT MALATE TRANSPORTER4 genotype determines malate content in cultivated jujube. Plant Physiology 191:414−427

doi: 10.1093/plphys/kiac491
[22]

Zhang C, Huang J, Li X. 2015. Identification of appropriate reference genes for RT-qPCR analysis in Ziziphus jujuba Mill. Scientia Horticulturae 197:166−169

doi: 10.1016/j.scienta.2015.09.026
[23]

Liu H, Zhao X, Bi J, Dong X, Zhang C. 2024. A natural mutation in the promoter of the aconitase gene ZjACO3 influences fruit citric acid content in jujube. Horticulture Research 11:uhae003

doi: 10.1093/hr/uhae003
[24]

Zhang Z, Shi Q, Wang B, Ma A, Wang Y, et al. 2022. Jujube metabolome selection determined the edible properties acquired during domestication. The Plant Journal 109:1116−1133

doi: 10.1111/tpj.15617
[25]

Huang J, Zhang C, Zhao X, Fei Z, Wan K, et al. 2016. The jujube genome provides insights into genome evolution and the domestication of sweetness/acidity taste in fruit trees. PLoS Genetics 12:e1006433

doi: 10.1371/journal.pgen.1006433
[26]

Dorn A, Bollekens J, Staub A, Benoist C, Mathis D. 1987. A multiplicity of CCAAT box-binding proteins. Cell 50:863−872

doi: 10.1016/0092-8674(87)90513-7
[27]

Wolucka BA, Persiau G, Van Doorsselaere J, Davey MW, Demol H, et al. 2001. Partial purification and identification of GDP-mannose 3', 5'-epimerase of Arabidopsis thaliana, a key enzyme of the plant vitamin C pathway. Proceedings of the National Academy of Sciences of the United States of America 98:14843−14848

doi: 10.1073/pnas.011578198
[28]

Ma L, Wang Y, Liu W, Liu Z. 2014. Overexpression of an alfalfa GDP-mannose 3, 5-epimerase gene enhances acid, drought and salt tolerance in transgenic Arabidopsis by increasing ascorbate accumulation. Biotechnology Letters 36:2331−2341

doi: 10.1007/s10529-014-1598-y
[29]

Liao G, Xu Q, Allan AC, Xu X. 2023. L-Ascorbic acid metabolism and regulation in fruit crops. Plant Physiology 192:1684−1695

doi: 10.1093/plphys/kiad241
[30]

van Huijsduijnen RAMH, Bollekens J, Dom A, Benoist C, Mathis D. 1987. Properties of a CCAAT box-binding protein. Nucleic Acids Research 15:7265−7282

doi: 10.1093/nar/15.18.7265
[31]

Pereira SLS, Martins CPS, Sousa AO, Camillo LR, Araújo CP, et al. 2018. Genome-wide characterization and expression analysis of citrus NUCLEAR FACTOR-Y (NF-Y) transcription factors identified a novel NF-YA gene involved in drought-stress response and tolerance. PLoS One 13:e0199187

doi: 10.1371/journal.pone.0199187
[32]

Li M, Du Q, Li J, Wang H, Xiao H, et al. 2023. Genome-wide identification and chilling stress analysis of the NF-Y gene family in melon. International Journal of Molecular Sciences 24:6934

doi: 10.3390/ijms24086934
[33]

Zhang Q, Zhang J, Wei H, Fu X, Ma L, et al. 2020. Genome-wide identification of NF-YA gene family in cotton and the positive role of GhNF-YA10 and GhNF-YA23 in salt tolerance. International Journal of Biological Macromolecules 165:2103−2115

doi: 10.1016/j.ijbiomac.2020.10.064
[34]

Wang T, Zou H, Ren S, Jin B, Lu Z. 2023. Genome-wide identification, characterization, and expression analysis of NF-Y gene family in Ginkgo biloba seedlings and GbNF-YA6 involved in heat-stress response and tolerance. International Journal of Molecular Sciences 24:12284

doi: 10.3390/ijms241512284
[35]

Chen M, Zhao Y, Zhuo C, Lu S, Guo Z. 2015. Overexpression of a NF-YC transcription factor from bermudagrass confers tolerance to drought and salinity in transgenic rice. Plant Biotechnology Journal 13:482−491

doi: 10.1111/pbi.12270
[36]

Ke X, Shen J, Niu Y, Zhao H, Guo Y, et al. 2023. Cucumber NUCLEAR FACTOR-YC2/-YC9 target translocon component CsTIC21 in chloroplast photomorphogenesis. Plant Physiology 192:2822−2837

doi: 10.1093/plphys/kiad296
[37]

Zhang C, Li W, Tan C, Huang M, Wu H, et al. 2025. Natural allelic variation in SW14 determines seed weight and quality in soybean. Nature Communications 16:8070

doi: 10.1038/s41467-025-63582-0
[38]

Laing WA, Martínez-Sánchez M, Wright MA, Bulley SM, Brewster D, et al. 2015. An upstream open reading frame is essential for feedback regulation of ascorbate biosynthesis in Arabidopsis. The Plant Cell 27:772−786

doi: 10.1105/tpc.114.133777