[1]

McGovern PE. 2003. Ancient wine: the search for the origins of viniculture. Princeton: Princeton University Press.

[2]

Myles S, Boyko AR, Owens CL, Brown PJ, Grassi F, et al. 2011. Genetic structure and domestication history of the grape. Proceedings of the National Academy of Sciences of the United States of America 108:3530−3535

doi: 10.1073/pnas.1009363108
[3]

McGovern P, Jalabadze M, Batiuk S, Callahan MP, Smith KE, et al. 2017. Early neolithic wine of Georgia in the South Caucasus. Proceedings of the National Academy of Sciences of the United States of America 114:E10309−E10318

doi: 10.1073/pnas.1714728114
[4]

Food and Agriculture Organization of the United Nations (FAO). 2023. FAO statistical databases. www.fao.org/faostat/zh/#data (accessed on 27 October 2025)

[5]

Sabra A, Netticadan T, Wijekoon C. 2021. Grape bioactive molecules, and the potential health benefits in reducing the risk of heart diseases. Food Chemistry: X 12:100149

doi: 10.1016/j.fochx.2021.100149
[6]

Oli KP, Pandey MR. 2024. The horizon of the third Pole: mapping future scenarios and strategic responses. Environmental Policy and Law 54:266−275

doi: 10.1177/18785395241293282
[7]

Xinhua. 2025. China's Tibet maintains stable ecological environment in 2024. https://english.news.cn/20250605/95db4ab070da4303bad087ba37b65bee/c.html (accessed on 27 September 2025)

[8]

Zhou T, Zhang W. 2021. Anthropogenic warming of Tibetan Plateau and constrained future projection. Environmental Research Letters 16:044039

doi: 10.1088/1748-9326/abede8
[9]

Zhu L, Ju J, Qiao B, Liu C, Wang J, et al. 2025. Physical and biogeochemical responses of Tibetan Plateau lakes to climate change. Nature Reviews Earth & Environment 6:284−298

doi: 10.1038/s43017-025-00650-5
[10]

Shen M, Wang S, Jiang N, Sun J, Cao R, et al. 2022. Plant phenology changes and drivers on the Qinghai-Tibetan Plateau. Nature Reviews Earth & Environment 3:633−651

doi: 10.1038/s43017-022-00317-5
[11]

Lin Z, Wu X. 1981. Climate zoning of the Qinghai-Tibet Plateau. Acta Geographica Sinica 36:22−32 (in Chinese)

[12]

International Organization of Vine and Wine. 2012. OIV Guidelines for vitiviniculture zoning methodologies on a soil and climate level. www.oiv.int/node/3271 (accessed on 20 April 2026)

[13]

Yang FS, Peng D, Ma XT, Ban Q, Du YF, et al. 2025. A review of plant diversity surveys and monographic studies of the Pan-Himalaya region. Journal of Systematics and Evolution 63:5−11 (in Chinese)

doi: 10.1111/jse.13161
[14]

Yang X, Zhuoga, Luobo. 2011. An overview of climate change and vegetation growth in Northern Tibet. Pratacultural Science 28:626−630 (in Chinese)

[15]

Zuo L, Dai A, Liu Q, Zuo C. 2004. Tibet the fruit tree resources rows with area the place explores. Tibet Journal of Agricultural Sciences 26:13−26 (in Chinese)

doi: 10.3969/j.issn.1005-2925.2004.03.004
[16]

Guge Q. 2013. Geography of Tibet. Beijing: Beijing Normal University Press. pp. 167−168

[17]

d'Alpoim Guedes J, Lu H, Li Y, Spengler RN, Wu X, et al. 2014. Moving agriculture onto the Tibetan Plateau: the archaeobotanical evidence. Archaeological and Anthropological Sciences 6:255−269

doi: 10.1007/s12520-013-0153-4
[18]

Han ISK. 2018. The journey to the east: the motif of grapes and grapevines along the silk roads. Acta Via Serica 3:107−134

[19]

Jiacuodunzhu. 1991. Tibetan murals: ethnic origins and political-religious history. Tibetan Art Studies 1991:80−91 (in Chinese)

[20]

Mao J. 2016. The mother Buddha herbal. Xining: Qinghai People's Publishing House. pp. 75

[21]

Zhu J. 1992. A mirror of auspicious animal grapes found in Bainang, Tibet. Cultural Relics 1992:8 (in Chinese)

[22]

Silangzhuoga. 2020. Rural revitalization strategy under the vision of economic research in YanJing, Tibet. Thesis. Tibet University, China. pp. 2

[23]

People's Government of Zogong County. 2025. Centennial old grape trees found in Zogong county, Tibet, with the maximum age of more than 400 years. https://zuogong.changdu.gov.cn/zgx/c105702/202506/b63cbb18c3064233bf467f2e1f1ddc92.shtml (accessed on 8 October 2025)

[24]

Wessels C. 1905. Antonio De Andrade. In Early Jesuit Travellers in Central Asia, 1603−1721. Dordrecht: Springer. pp. 43–68 doi: 10.1007/978-94-017-6836-8_2

[25]

Galipeau BA. 2017. Tibetan wine production, taste of place, and regional niche identities in Shangri-La, China. In Trans-Himalayan borderlands: Livelihoods, territorialities, modernities, eds Yü DS, Michaud J. Amsterdam: Amsterdam University Press. pp. 207−228

[26]

Salick J, Byg A, Konchar K. 2011. Innovation and adaptation in Tibetan land use and agriculture coping with climate change. In Indigenous Peoples, Marginalized Populations and Climate Change: Vulnerability, Adaptation and Traditional Knowledge. Proceedings of the Conference Indigenous People, Marginalized Populations and Climate Change. Mexico City, Mexico, 2011. pp. 1−14

[27]

Duan S, Zong X, Liu X, Zuo Y, Duan Y. 1983. Preliminary report on the fruit germplasm resources in the Tibet Autonomous Region. Acta Horticulturae Sinica 10:217−223 (in Chinese)

[28]

Zong X, Duan Y, Zuo Y. 1983. Fruit tree resources in Qamdo, Tibet. China Seed Industry 1983:25−33 (in Chinese)

[29]

Zhang H, Li B. 2012. Current status and development strategies of protected horticultural fruit trees in the Tibetan Plateau. South China Agriculture 6:68−70 (in Chinese)

[30]

Zhang X, Xing S, Song X, Wang L. 2019. Analysis on the current situation of grape cultivation in Lhasa area and its countermeasures. Heilongjiang Agricultural Sciences 2019:132−134 (in Chinese)

[31]

Cai S, Cai Z, Lin Y, Yu D. 2009. Introduction and related cultivation technology of Vitis vinifera in the Tibetan Plateau. Fujian Fruits 2009:61−63 (in Chinese)

[32]

Zhang L. 2016. Losong Ciren crafts wine on the roof of the world. New West 2016:71−75 (in Chinese)

[33]

Shen Y. 2019. Vineyard report 2019 (Issue 18): summer in Tibet - the Pristine Blue Highlands. www.winesou.com/news/china_news/140454.html (accessed on 24 April 2026)

[34]

Tibet Academy of Agricultural and Animal Husbandry Sciences (TAAAS). 2025. The characteristic grape industry in Tibet has become another significant channel for increasing income and prosperity among farmers and herdsmen. www.taaas.org/info/1003/14691.htm (accessed on 16 September 2025)

[35]

Shi X, Li Y, Jiang S, Xu G, Huang J, et al. 2025. Current status and optimization strategies for the fruit industry in Tibet. China Fruits 2025:134−138 (in Chinese)

[36]

Lu G. 2019. Preliminary study on secondary results of Lhasa facility grape. Tibet Agricultural Science and Technology 41:36−39 (in Chinese)

doi: 10.3969/j.issn.1005-2925.2019.01.011
[37]

National Bureau of Statistics of China. 2024. China Statistical yearbook 2023. www.stats.gov.cn/sj/ndsj/2024/indexch.htm (accessed on 18 October 2025)

[38]

Protected Geographical Indication (PGI). 2017. Wine fragrance from the highlands: chronicles of Yanjing wine, China's PGI product. Standard Living 2017:92−96 (in Chinese)

[39]

Huebra LJF, de Araujo Gastal S. 2023. Os vinhos de altura em Salta, Argentina: um estudo sobre as bodegas Colomé e Domigo Hermanos. Revista Memória em Rede 15:315−318

doi: 10.15210/rmr.v15i28.23916
[40]

Luo R, Zhang W, Li Y, Bai M, Lu X, et al. 2024. Breeding report of a new wine grape variety Yunniang 3. Journal of Fruit Science 41:2130−2133 (in Chinese)

doi: 10.13925/j.cnki.gsxb.20240332
[41]

Deng L, Tan H, Jiang B, Wang D, Sun L, et al. 2025. Investigation and conservation of wild berry and other germplasm resources in Tibet. Xizang Science and Technology 47:10−21,40 (in Chinese)

doi: 10.3969/j.issn.1004-3403.2025.07.003
[42]

Li G, Xie Z, Zheng Y, Chen W, Cai S, et al. 2008. Countermeasures of fruit industry sustainable development in Tibet. Acta Horticulturae Sinica 35:899−908 (in Chinese)

doi: 10.3321/j.issn:0513-353X.2008.06.021
[43]

Li Y, Zhou Y, Liu F, Liu X, Wang Q. 2022. Diversity patterns of wetland angiosperms in the Qinghai-Tibet Plateau, China. Diversity 14:777

doi: 10.3390/d14100777
[44]

Duan S, Zong X, Duan Y. 1984. Species and geographical distribution of fruit trees in Tibet. China Fruits 1984:20−26 (in Chinese)

[45]

Wan Y, Schwaninger H, Li D, Simon CJ, Wang Y, et al. 2008. The eco-geographic distribution of wild grape germplasm in China. Vitis 47:77−80

[46]

Zheng H, Li J, Wang S, Zheng Y. 2004. Records of fruit germplasm resources in Tibet. Beijing: China Agriculture Press. pp. 125−136

[47]

You Y, Yu J, Nie Z, Peng D, Barrett RL, et al. 2024. Transition of survival strategies under global climate shifts in the grape family. Nature Plants 10:1100−1111

doi: 10.1038/s41477-024-01726-8
[48]

Liang F. 2021. Key cultivation techniques of seedless white-heart grape in greenhouse in Ali region of Tibet. Tibet Journal of Agricultural Sciences 43:65−67 (in Chinese)

doi: 10.3969/j.issn.1005-2925.2021.04.017
[49]

Pan G. 1995. Protected cultivation techniques for Kyoho grapes in Tibet. Forest By-Product and Speciality in China 1995:18−19 (in Chinese)

[50]

Li Y. 2016. Lhasa introduction of new varieties of grape experimental research. Tibet Journal of Agricultural Sciences 38:13−17 (in Chinese)

doi: 10.3969/j.issn.1005-2925.2016.03.003
[51]

Zhu Z, Yang R. 2024. Introduction performance and key cultivation techniques of Barbera grapes in solar greenhouses of Qushui, Tibet. Northwest Horticulture 2024:38−40,83 (in Chinese)

[52]

Zhang K, Cao J, Wang W, Wang J, Hui Z. 2023. Grape quality differences at different altitudes in Shangri-La, Yunnan. Journal of Northwest A& F University (Natural Science Edition) 51:129−137 (in Chinese)

[53]

Mansour G, Ghanem C, Mercenaro L, Nassif N, Hassoun G, et al. 2022. Effects of altitude on the chemical composition of grapes and wine: a review. OENO One 56:227−239

doi: 10.20870/oeno-one.2022.56.1.4895
[54]

Bai S, Tao X, Hu J, Chen H, Wu J, et al. 2025. Flavonoids profile and antioxidant capacity of four wine grape cultivars and their wines grown in the Turpan Basin of China, the hottest wine region in the world. Food Chemistry: X 26:102301

doi: 10.1016/j.fochx.2025.102301
[55]

Wu Y, Ji C, Jiang Y, Hu H, Yu T, et al. 2024. Mechanisms of Meyerozyma caribbica isolated from Tibetan soil to inhibit Aspergillus ochraceus on grapes. Postharvest Biology and Technology 210:112797

doi: 10.1016/j.postharvbio.2024.112797
[56]

Li L, Zhou Y, Tsoja W, Norsong G, Lagba T, et al. 2025. Spatiotemporal variations of solar spectrum under different altitudes and latitudes on the Qinghai-Tibet plateau. Acta Optica Sinica 2:44−51 (in Chinese)

[57]

Martínez-Lüscher J, Sánchez-Díaz M, Delrot S, Aguirreolea J, Pascual I, et al. 2014. Ultraviolet-B radiation and water deficit interact to alter flavonol and anthocyanin profiles in grapevine berries through transcriptomic regulation. Plant and Cell Physiology 55:1925−1936

doi: 10.1093/pcp/pcu121
[58]

Liang T, Shi C, Peng Y, Tan H, Xin P, et al. 2020. Brassinosteroid-activated BRI1-EMS-SUPPRESSOR 1 inhibits flavonoid biosynthesis and coordinates growth and UV-B stress responses in plants. The Plant Cell 32:3224−3239

doi: 10.1105/tpc.20.00048
[59]

Liang J, Guo J, Liu Y, Zhang Z, Zhou R, et al. 2023. UV-C promotes the accumulation of flavane-3-ols in juvenile fruit of grape through positive regulating VvMYBPA1. Plants 12:1691

doi: 10.3390/plants12081691
[60]

Shao J, Ma C, Liang X, Du L. 2011. Measurement and analysis of the content of total ployphenols and proanthocyanidins in Carbernet Sauvignon dry red wine from Yun'nan and from other regions. Liquor-Making Science & Technology 2011:29−32 (in Chinese)

[61]

Zhang K, Yin H, Wang J, Cao J, Xi Z. 2023. Differences and genesis of grape phenolic compounds among different altitudes in Yunnan Shangri-la. Scientia Agricultura Sinica 56:3879−3893 (in Chinese)

doi: 10.3864/j.issn.0578-1752.2023.19.014
[62]

Jiang B, Fu F, Sun Z, Wang L. 2016. Research progress on influence of cultivation altitude on phenolics of grape berry and wine. Food and Fermentation Industries 42:262−267 (in Chinese)

doi: 10.13995/j.cnki.11-1802/ts.201608044
[63]

Jin XD, Wu X, Liu X. 2017. Phenolic characteristics and antioxidant activity of merlot and cabernet sauvignon wines increase with vineyard altitude in a high-altitude region. South African Journal of Enology & Viticulture 38:132−143

doi: 10.21548/38-2-1068
[64]

Mao R, Zhang G, Wang J, Du F, Deng W, et al. 2015. Shiraz/Syrah grape metabolites response to low/high altitudes. Modern Agricultural Science and Technology 2015:240−246 (in Chinese)

doi: 10.3969/j.issn.1007-5739.2015.20.134
[65]

Mao R, Zhang G, Shao J, Du F, Deng W, et al. 2016. Response of 'Merlot' grape berry quality and metabolome to meteorological factors at both low and high altitudes. Chinese Journal of Eco-Agriculture 24:506−516 (in Chinese)

[66]

Hu X. 2025. Study on terroir characteristics of vineyards at different altitudes in high mountain area of Southwest China. Thesis. Northwest A&F University, China. pp. 43−85

[67]

Yin H, Wang L, Wu Y, Xi Z. 2024. Roles of non-visible light and temperature in the regulation of anthocyanin synthesis in fruits and vegetables. Food Frontiers 5:1968−1983

doi: 10.1002/fft2.426
[68]

Zheng X, Graburn N. 2022. Terroir aura: Tibetan wine as gift in China's southwest. In Wine and the gift. London: Routledge. pp. 90−107 doi: 10.4324/9781003038986-6

[69]

Wang Y, Wang L, Liu X, Li Y, Wang X, et al. 2018. Climatic regionalization of wine grapes in the Hengduan Mountain region of China. Spanish Journal of Agricultural Research 16:e0303

doi: 10.5424/sjar/2018162-12457
[70]

Liu J. 2016. Research on Himalayan region wine industrial cluster innovation and management. In Proceedings of the 2016 China Wine Forum Yantai. Yantai: China Alcoholic Drinks Association. pp. 252−269

[71]

Jiu X, Zhan D. 2023. The classification of Tibetan wine and its health benefit. International Journal of Food Science and Agriculture 7:285−289

doi: 10.26855/ijfsa.2023.06.018
[72]

Kan J, Wu L, Ren H, Zhong Z. 2019. Fermentation technique optimization of wine with two grape varieties at natural temperature. Journal of Plateau Agriculture 3:94−102 (in Chinese)

doi: 10.19707/j.cnki.jpa.2019.01.016
[73]

Sun C, Xing Y, Jiang L, Li L, Dong Z, et al. 2015. Effect of table grape raw material on the quality of ice grape wine in plateau of Sichuan province. The Food Industry 36:95−99 (in Chinese)

[74]

Du C, Zhang K, Lin Q, Huang S, Han Y, et al. 2025. Rapid ecological change outpaces climate warming in Tibetan glacier lakes. Communications Earth & Environment 6:523

doi: 10.1038/s43247-025-02443-6
[75]

Sun C, Xu XD, Wang PJ, Yang D, Zhang SJ, et al. 2022. The warming and wetting ecological environment changes over the Qinghai-Tibetan Plateau and the driving effect of the Asian summer monsoon. Journal of Tropical Meteorology 28:95−108

doi: 10.46267/j.1006-8775.2022.008
[76]

Liu X, Chen B. 2000. Climatic warming in the Tibetan Plateau during recent decades. International Journal of Climatology 20:1729−1742

doi: 10.1002/1097-0088(20001130)20:14<1729::AID-JOC556>3.0.CO;2-Y
[77]

Liu X, Zhou Q, Ma Y, Zhi Z, Liu R, et al. 2025. Temporal and spatial variation characteristics of precipitation isohyets on the Qinghai–Tibet Plateau from 1961 to 2023. Atmosphere 16:698

doi: 10.3390/atmos16060698
[78]

Li Y, Hou Z, Zhang L, Song C, Piao S, et al. 2023. Rapid expansion of wetlands on the Central Tibetan Plateau by global warming and El Niño. Science Bulletin 68:485−488

doi: 10.1016/j.scib.2023.02.021
[79]

Wei X, Zhang Y, Sun S, Song Z. 2025. The response characteristics of vegetation to climate change of the growing season on the Tibetan Plateau. Earth and Environment 53:631−642

doi: 10.3724/EE.1672-9250.2025.53.033
[80]

Lamchin M, Lee WK, Wang SW. 2022. Multi-temporal analysis of past and future land-cover changes of the third pole. Land 11:2227

doi: 10.3390/land11122227
[81]

Van Leeuwen C, Sgubin G, Bois B, Ollat N, Swingedouw D, et al. 2024. Climate change impacts and adaptations of wine production. Nature Reviews Earth & Environment 5:258−275

doi: 10.1038/s43017-024-00521-5
[82]

Liu Y, Shi X, Du H, Jiang M, Li F, et al. 2025. Spatiotemporal distribution of wine grape under climate change in Northwestern China. Ecology and Evolution 15:e70826

doi: 10.1002/ece3.70826
[83]

Arias LA, Berli F, Fontana A, Bottini R, Piccoli P. 2022. Climate change effects on grapevine physiology and biochemistry: benefits and challenges of high altitude as an adaptation strategy. Frontiers in Plant Science 13:835425

doi: 10.3389/fpls.2022.835425
[84]

Zhang K, Cao J, Yin H, Wang J, Wang X, et al. 2024. Microclimate diversity drives grape quality difference at high-altitude: observation using PCA analysis and structural equation modeling (SEM). Food Research International 191:114644

doi: 10.1016/j.foodres.2024.114644
[85]

Buytaert W, Cuesta-Camacho F, Tobón C. 2011. Potential impacts of climate change on the environmental services of humid tropical Alpine regions. Global Ecology and Biogeography 20:19−33

doi: 10.1111/j.1466-8238.2010.00585.x
[86]

Martínez-Lüscher J, Sánchez-Díaz M, Delrot S, Aguirreolea J, Pascual I, et al. 2016. Ultraviolet-B alleviates the uncoupling effect of elevated CO2 and increased temperature on grape berry (Vitis vinifera cv. Tempranillo) anthocyanin and sugar accumulation: effect of UV-B, elevated CO2 and increased temperature. Australian Journal of Grape and Wine Research 22:87−95

doi: 10.1111/ajgw.12213
[87]

Van Leeuwen C, Destrac-Irvine A. 2017. Modified grape composition under climate change conditions requires adaptations in the vineyard. OENO One 51:147−154

doi: 10.20870/oeno-one.2016.0.0.1647
[88]

Cataldo E, Eichmeier A, Mattii GB. 2023. Effects of global warming on grapevine berries phenolic compounds — a review. Agronomy 13:2192

doi: 10.3390/agronomy13092192
[89]

Gambetta JM, Holzapfel BP, Stoll M, Friedel M. 2021. Sunburn in grapes: a review. Frontiers in Plant Science 11:604691

doi: 10.3389/fpls.2020.604691
[90]

Muntean MD, Tomoiagă LL, Răcoare HS, Sîrbu AD, Giurcă IS, et al. 2025. Millerandage — one of the grapevine cultivation challenges in the climate change context. Horticulturae 11:165

doi: 10.3390/horticulturae11020165
[91]

Zhou Y, Yuan G, Cong Z, Wang X. 2021. Priorities for the sustainable development of the ecological environment on the Tibetan Plateau. Fundamental Research 1:329−333

doi: 10.1016/j.fmre.2021.03.006
[92]

Ding Z, Ha Y, Hu Y, Zhu Y, Dai H, et al. 2025. Spatiotemporal characteristics of summer extreme precipitation over the Inner Tibetan Plateau in recent decades. npj Climate and Atmospheric Science 8:193

doi: 10.1038/s41612-025-01089-7
[93]

Alikadic A, Pertot I, Eccel E, Dolci C, Zarbo C, et al. 2019. The impact of climate change on grapevine phenology and the influence of altitude: a regional study. Agricultural and Forest Meteorology 271:73−82

doi: 10.1016/j.agrformet.2019.02.030
[94]

Li X, Duan A, Shangguan D, Wang L, Li C, et al. 2025. Future projection of climate change in the three poles. In The Three Poles of the Earth: Challenges to Sustainable Development in Fragile Environments. Singapore: Springer. pp. 27−73 doi: 10.1007/978-981-97-7721-1_2

[95]

Zhang K, Wang S, Lu G, Tudan J, Li Y. 2019. Effect of modified-atmosphere packaging on quality of 'Red Earth' grape during storage in Lhasa. Tibet Journal of Agricultural Sciences 41:17−20 (in Chinese)

[96]

Lu G, Zhang K, Min Z, Zhao H, Tudan J, et al. 2019. Key points of cultivation and management of facility grape in Tibet. China Fruit & Vegetable 39:84−87 (in Chinese)

doi: 10.19590/j.cnki.1008-1038.2019.09.023
[97]

Zhang K, Zhou J, Zhao L. 2024. Application progress of pruning in grape production. Tibet Journal of Agricultural Sciences 46:71−74 (in Chinese)

doi: 10.3969/j.issn.1005-2925.2024.02.017
[98]

Zhao L, Zhou J, Zhang K. 2024. High-yield cultivation technology of Red Globe grape in facilities in Lhasa. Tibet Journal of Agricultural Sciences 46:59−61 (in Chinese)

doi: 10.3969/j.issn.1005-2925.2024.03.014
[99]

Yang R, Zhu Z, Guo J, Qin B. 2022. Investigation on occurrence of pests and diseases in facility-grown grapes in Lhasa surrounding areas. Contemporary Horticulture 45:84−85,100 (in Chinese)

[100]

Wang S, Li S, Zeng X, Dai A, Zhang S, et al. 2018. Soil acidification, salinization and nutrient characteristics in greenhouse vineyards in Tibet. Journal of Fruit Science 35:957−966 (in Chinese)