[1]

Yang ZY, Zhao LY, Xu ZD. 2010. 野生玫瑰与栽培玫瑰对盐胁迫反应的比较研究 [Comparison of the resistance to salt stress between wild plants and cultivars of Rosa rugosa]. 山东林业科技 [Shandong Forestry Science and Technology] 40:43−44,50 (in Chinese)

doi: 10.3969/j.issn.1002-2724.2010.02.013
[2]

Chen F, Su L, Hu S, Xue JY, Liu H, et al. 2021. A chromosome-level genome assembly of rugged rose (Rosa rugosa) provides insights into its evolution, ecology, and floral characteristics. Horticulture Research 8:141

doi: 10.1038/s41438-021-00594-z
[3]

Patel S. 2017. Rose hip as an underutilized functional food: evidence-based review. Trends in Food Science & Technology 63:29−38

doi: 10.1016/j.tifs.2017.03.001
[4]

Cendrowski A, Kraśniewska K, Przybył JL, Zielińska A, Kalisz S. 2020. Antibacterial and antioxidant activity of extracts from rose fruits (Rosa rugosa). Molecules 25:1365

doi: 10.3390/molecules25061365
[5]

Çolak AM , Alan F. 2025. Determination of pomological and chemical properties of some rosehip (Rosa spp.) genotypes growing naturally in Kayseri province. ISPEC Journal of Agricultural Sciences 9:621−629

doi: 10.5281/zenodo.15509973
[6]

Duarte M, Santos Pedrosa S, Khusial PR, Madureira AR. 2025. The biological potential and health-benefits of flavonoids: a review and development opportunities. Chemico-Biological Interactions 421:111755

doi: 10.1016/j.cbi.2025.111755
[7]

Peluso I, Miglio C, Morabito G, Ioannone F, Serafini M. 2015. Flavonoids and immune function in human: a systematic review. Critical Reviews in Food Science and Nutrition 55:383−395

doi: 10.1080/10408398.2012.656770
[8]

Sarma AD, Sreelakshmi Y, Sharma R. 1997. Antioxidant ability of anthocyanins against ascorbic acid oxidation. Phytochemistry 45:671−674

doi: 10.1016/S0031-9422(97)00057-5
[9]

Yao LH, Jiang YM, Shi J, Tomás-Barberán FA, Datta N, et al. 2004. Flavonoids in food and their health benefits. Plant Foods for Human Nutrition 59:113−122

doi: 10.1007/s11130-004-0049-7
[10]

Dolek U, Gunes M, Genc N, Elmastas M. 2018. Total phenolic compound and antioxidant activity changes in rosehip (Rosa sp.) during ripening. Journal of Agricultural Science and Technology 20:817−828

[11]

Andersson SC, Rumpunen K, Johansson E, Olsson ME. 2011. Carotenoid content and composition in rose hips (Rosa spp.) during ripening, determination of suitable maturity marker and implications for health promoting food products. Food Chemistry 128:689−696

doi: 10.1016/j.foodchem.2011.03.088
[12]

Medveckienė B, Levickienė D, Vaitkevičienė N, Vaštakaitė-Kairienė V, Kulaitienė J. 2023. Changes in pomological and physical parameters in rosehips during ripening. Plants 12:1314

doi: 10.3390/plants12061314
[13]

Arora N, Lo E, Philippidis GP. 2022. A two-prong mutagenesis and adaptive evolution strategy to enhance the temperature tolerance and productivity of Nannochloropsis oculata. Bioresource Technology 364:128101

doi: 10.1016/j.biortech.2022.128101
[14]

Appelhagen I, Lu GH, Huep G, Schmelzer E, Weisshaar B, et al. 2011. TRANSPARENT TESTA1 interacts with R2R3-MYB factors and affects early and late steps of flavonoid biosynthesis in the endothelium of Arabidopsis thaliana seeds. The Plant Journal 67:406−419

doi: 10.1111/j.1365-313X.2011.04603.x
[15]

Liu S, Zhang H, Meng Z, Jia Z, Fu F, et al. 2025. The LncNAT11–MYB11–F3'H/FLS module mediates flavonol biosynthesis to regulate salt stress tolerance in Ginkgo biloba. Journal of Experimental Botany 76:1179−1201

doi: 10.1093/jxb/erae438
[16]

Feller A, Machemer K, Braun EL, Grotewold E. 2011. Evolutionary and comparative analysis of MYB and bHLH plant transcription factors. The Plant Journal 66:94−116

doi: 10.1111/j.1365-313X.2010.04459.x
[17]

Shi Y , Lu T , Lai S, Li S, Zhang L, et al. 2024. Rosa rugosa R2R3-MYB transcription factors RrMYB12 and RrMYB111 regulate the accumulation of flavonols and anthocyanins. Frontiers in Plant Science 15:1477278

doi: 10.3389/fpls.2024.1477278
[18]

Chen NF, Zhang L. 2005. 金樱子黄酮类化合物的初步研究 [Preliminary study on flavonoid compounds from Rosa laevigata Michx]. 中国林副特产 [Forest By-Product and Speciality in China] 2005:2−4 (in Chinese)

doi: 10.13268/j.cnki.fbsic.2005.05.002
[19]

Wiseman H, O'Reilly JD, Adlercreutz H, Mallet AI, Bowey EA, et al. 2000. Isoflavone phytoestrogens consumed in soy decrease F2-isoprostane concentrations and increase resistance of low-density lipoprotein to oxidation in humans. The American Journal of Clinical Nutrition 72:395−400

doi: 10.1093/ajcn/72.2.395
[20]

Erlund I, Kosonen T, Alfthan G, Mäenpää J, Perttunen K, et al. 2000. Pharmacokinetics of quercetin from quercetin aglycone and rutin in healthy volunteers. European Journal of Clinical Pharmacology 56:545−553

doi: 10.1007/s002280000197
[21]

Vyniarska AV. 2024. Quercetin: biological activity, therapeutic potential and prospects of the use. Scientific Messenger of LNU of Veterinary Medicine and Biotechnologies 26:242−247

doi: 10.32718/nvlvet11635
[22]

Adhimi CS, Sakti AS, Pratiwi ED, Khan MA. 2024. Article review: in vitro, in vivo, and clinical trial data on proanthocyanidin compounds. Surya 16:63−71

doi: 10.38040/js.v16i2.1011
[23]

Liu C, Bolling BW. 2024. Dietary proanthocyanidins for improving gut immune health. Current Opinion in Food Science 56:101133

doi: 10.1016/j.cofs.2024.101133
[24]

Lücker J, Martens S, Lund ST. 2010. Characterization of a Vitis vinifera cv. Cabernet Sauvignon 3', 5'-O-methyltransferase showing strong preference for anthocyanins and glycosylated flavonols. Phytochemistry 71:1474−1484

doi: 10.1016/j.phytochem.2010.05.027
[25]

Fournier-Level A, Hugueney P, Verriès C, This P, Ageorges A. 2011. Genetic mechanisms underlying the methylation level of anthocyanins in grape (Vitis vinifera L.). BMC Plant Biology 11:179

doi: 10.1186/1471-2229-11-179
[26]

D'Auria JC, Reichelt M, Luck K, Svatoš A, Gershenzon J. 2007. Identification and characterization of the BAHD acyltransferase malonyl CoA: anthocyanidin 5-O-glucoside-6''-O-malonyltransferase (At5MAT) in Arabidopsis thaliana. FEBS Letters 581:872−878

doi: 10.1016/j.febslet.2007.01.060
[27]

Suzuki H, Nakayama T, Nishino T. 2003. Proposed mechanism and functional amino acid residues of malonyl-CoA: anthocyanin 5-O-glucoside-6'''-O-malonyltransferase from flowers of Salvia splendens, a member of the versatile plant acyltransferase family. Biochemistry 42:1764−1771

doi: 10.1021/bi020618g
[28]

Quattrocchio F, Verweij W, Kroon A, Spelt C, Mol J, et al. 2006. PH4 of Petunia is an R2R3 MYB protein that activates vacuolar acidification through interactions with basic-helix-loop-helix transcription factors of the anthocyanin pathway. The Plant Cell 18:1274−1291

doi: 10.1105/tpc.105.034041
[29]

Elomaa P, Uimari A, Mehto M, Albert VA, Laitinen RAE, et al. 2003. Activation of anthocyanin biosynthesis in Gerbera hybrida (Asteraceae) suggests conserved protein-protein and protein promoter interactions between the anciently diverged monocots and eudicots. Plant Physiology 133:1831−1842

doi: 10.1104/pp.103.026039
[30]

Mathews H, Clendennen SK, Caldwell CG, Liu XL, Connors K, et al. 2003. Activation tagging in tomato identifies a transcriptional regulator of anthocyanin biosynthesis, modification, and transport. The Plant Cell 15:1689−1703

doi: 10.1105/tpc.012963
[31]

Kobayashi S, Goto-Yamamoto N, Hirochika H. 2004. Retrotransposon-induced mutations in grape skin color. Science 304:982

doi: 10.1126/science.1095011
[32]

Espley RV, Hellens RP, Putterill J, Stevenson DE, Kutty-Amma S, et al. 2007. Red colouration in apple fruit is due to the activity of the MYB transcription factor, MdMYB10. The Plant Journal 49:414−427

doi: 10.1111/j.1365-313X.2006.02964.x
[33]

Nakatsuka T, Haruta KS, Pitaksutheepong C, Abe Y, Kakizaki Y, et al. 2008. Identification and characterization of R2R3-MYB and bHLH transcription factors regulating anthocyanin biosynthesis in gentian flowers. Plant and Cell Physiology 49:1818−1829

doi: 10.1093/pcp/pcn163
[34]

Schwinn K, Venail J, Shang Y, Mackay S, Alm V, et al. 2006. A small family of MYB-regulatory genes controls floral pigmentation intensity and patterning in the genus Antirrhinum. The Plant Cell 18:831−851

doi: 10.1105/tpc.105.039255
[35]

Nesi N, Jond C, Debeaujon I, Caboche M, Lepiniec L. 2001. The Arabidopsis TT2 gene encodes an R2R3 MYB domain protein that acts as a key determinant for proanthocyanidin accumulation in developing seed. The Plant Cell 13:2099−2114

doi: 10.1105/tpc.010098
[36]

Spelt C, Quattrocchio F, Mol JNM, Koes R. 2000. Anthocyanin1 of petunia encodes a basic helix-loop-helix protein that directly activates transcription of structural anthocyanin genes. The Plant Cell 12:1619−1631

doi: 10.1105/tpc.12.9.1619