[1]

Zhang X, Wu Q, Zhao Y, Aimy A, Yang X. 2019. Consumption of post-fermented Jing-Wei Fuzhuan brick tea alleviates liver dysfunction and intestinal microbiota dysbiosis in high fructose diet-fed mice. RSC Advances 9:17501−17513

doi: 10.1039/C9RA02473E
[2]

Chen H, Ma Z, Xu Z. 2024. The tea road and the tea culture heritage of Hunan Province. In Cultural Heritage Preservation for Vulnerable Territories, eds Augelli F, Rigamonti M. Vol 6. Cham: Springer. pp. 17–40 doi: 10.1007/978-3-031-54390-6_2
[3]

Feng X, Chen M, Song H, Ma S, Ou C, et al. 2023. A systemic review on Liubao tea: a time-honored dark tea with distinctive raw materials, process techniques, chemical profiles, and biological activities. Comprehensive Reviews in Food Science and Food Safety 22:5063−5085

doi: 10.1111/1541-4337.13254
[4]

Li Q, Huang J, Li Y, Zhang Y, Luo Y, et al. 2017. Fungal community succession and major components change during manufacturing process of Fu brick tea. Scientific Reports 7:6947

doi: 10.1038/s41598-017-07098-8
[5]

Huang X, Li Y, Zhou F, Xiao T, Shang B, et al. 2024. Insight into the chemical compositions of Anhua dark teas derived from identical tea materials: a multi-omics, electronic sensory, and microbial sequencing analysis. Food Chemistry 441:138367

doi: 10.1016/j.foodchem.2024.138367
[6]

Zhang L, Zhang ZZ, Zhou YB, Ling TJ, Wan XC. 2013. Chinese dark teas: postfermentation, chemistry and biological activities. Food Research International 53:600−607

doi: 10.1016/j.foodres.2013.01.016
[7]

Shi J, Ma W, Wang C, Wu W, Tian J, et al. 2021. Impact of various microbial-fermented methods on the chemical profile of dark tea using a single raw tea material. Journal of Agricultural and Food Chemistry 69:4210−4222

doi: 10.1021/acs.jafc.1c00598
[8]

Liu Y, Huang W, Zhang C, Li C, Fang Z, et al. 2022. Targeted and untargeted metabolomic analyses and biological activity of Tibetan tea. Food Chemistry 384:132517

doi: 10.1016/j.foodchem.2022.132517
[9]

Ning J, Li D, Luo X, Ding D, Song Y, et al. 2016. Stepwise identification of six tea (Camellia sinensis (L.)) categories based on catechins, caffeine, and theanine contents combined with fisher discriminant analysis. Food Analytical Methods 9:3242−3250

doi: 10.1007/s12161-016-0518-2
[10]

Lv HP, Zhang Y, Shi J, Lin Z. 2017. Phytochemical profiles and antioxidant activities of Chinese dark teas obtained by different processing technologies. Food Research International 100:486−493

doi: 10.1016/j.foodres.2016.10.024
[11]

An T, Chen M, Zu Z, Chen Q, Lu H, et al. 2021. Untargeted and targeted metabolomics reveal changes in the chemical constituents of instant dark tea during liquid-state fermentation by Eurotium cristatum. Food Research International 148:110623

doi: 10.1016/j.foodres.2021.110623
[12]

Xiao Y, Li M, Wu Y, Zhong K, Gao H. 2020. Structural characteristics and hypolipidemic activity of theabrownins from dark tea fermented by single species Eurotium cristatum PW-1. Biomolecules 10:204

doi: 10.3390/biom10020204
[13]

Cheng L, Wei Y, Peng L, Wei K, Liu Z, et al. 2023. State-of-the-art review of theabrownins: from preparation, structural characterization to health-promoting benefits. Critical Reviews in Food Science and Nutrition 64:11321−11340

doi: 10.1080/10408398.2023.2236701
[14]

Liu C, Liao Y, Wu A, He C, Du X, et al. 2025. Tibetan dark tea Theabrownin alleviates LPS-induced inflammation by modulating the Nrf2/NF-κB signaling pathway and host microbial metabolites. Food Chemistry 483:144264

doi: 10.1016/j.foodchem.2025.144264
[15]

Wang W, Zhang L, Wang S, Shi S, Jiang Y, et al. 2014. 8-C N-ethyl-2-pyrrolidinone substituted flavan-3-ols as the marker compounds of Chinese dark teas formed in the post-fermentation process provide significant antioxidative activity. Food Chemistry 152:539−545

doi: 10.1016/j.foodchem.2013.10.117
[16]

Zhou ZH, Zhang YJ, Xu M, Yang CR. 2005. Puerins A and B, two new 8-C substituted flavan-3-ols from Pu-er tea. Journal of Agricultural and Food Chemistry 53:8614−8617

doi: 10.1021/jf051390h
[17]

Tao MK, Xu M, Zhu HT, Cheng RR, Wang D, et al. 2014. New phenylpropanoid-substituted flavan-3-ols from Pu-er ripe tea. Natural Product Communications 9:1167−1170

doi: 10.1177/1934578x1400900827
[18]

Tian LW, Tao MK, Xu M, Hu J, Zhu HT, et al. 2014. Carboxymethyl- and carboxyl-catechins from ripe Pu-er tea. Journal of Agricultural and Food Chemistry 62:12229−12234

doi: 10.1021/jf5036959
[19]

Li Q, Liu Z, Huang J, Luo G, Liang Q, et al. 2013. Anti-obesity and hypolipidemic effects of Fuzhuan brick tea water extract in high-fat diet-induced obese rats. Journal of the Science of Food and Agriculture 93:1310−1316

doi: 10.1002/jsfa.5887
[20]

Zhu MZ, Li N, Zhou F, Ouyang J, Lu DM, et al. 2020. Microbial bioconversion of the chemical components in dark tea. Food Chemistry 312:126043

doi: 10.1016/j.foodchem.2019.126043
[21]

Jiang HY, Shii T, Matsuo Y, Tanaka T, Jiang ZH, et al. 2011. A new catechin oxidation product and polymeric polyphenols of post-fermented tea. Food Chemistry 129:830−836

doi: 10.1016/j.foodchem.2011.05.031
[22]

Kanegae A, Sakamoto A, Nakayama H, Nakazono Y, Yakashiro I, et al. 2013. New phenolic compounds from Camellia sinensis L. fermented leaves. Journal of Natural Medicines 67:652−656

doi: 10.1007/s11418-012-0704-5
[23]

Chen M, Zhu Y, Zhang H, Wang J, Liu X, et al. 2017. Phenolic compounds and the biological effects of Pu-erh teas with long-term storage. International Journal of Food Properties 20:1715−1728

doi: 10.1080/10942912.2016.1217877
[24]

Lu H, Lin Z, Zhong Q, Wang L. 2010. Study on the chemical component of E8 fraction from Pu-erh tea. Journal of Tea Science 30:423−428
[25]

Wang S, Qiu Y, Gan RY, Zhu F. 2022. Chemical constituents and biological properties of Pu-erh tea. Food Research International 154:110899

doi: 10.1016/j.foodres.2021.110899
[26]

Luo ZM, Du HX, Li LX, An MQ, Zhang ZZ, et al. 2013. Fuzhuanins A and B: the B-ring fission lactones of flavan-3-ols from Fuzhuan brick-tea. Journal of Agricultural and Food Chemistry 61:6982−6990

doi: 10.1021/jf401724w
[27]

Zhang L, Li N, Ma ZZ, Tu PF. 2011. Comparison of the chemical constituents of aged Pu-erh tea, ripened Pu-erh tea, and other teas using HPLC-DAD-ESI-MSn. Journal of Agricultural and Food Chemistry 59:8754−8760

doi: 10.1021/jf2015733
[28]

She G, Chen K, Zhang Y, Yang C. 2007. The occurrence of 8-oxocaffeine and pyrimidine alkaloids in Pu-Er ripe tea. Acta Botanica Yunnanica 29:713−716

doi: 10.3969/j.issn.2095-0845.2007.06.019
[29]

Tao MK, Xu M, Zhang H, Chen H, Liu C, et al. 2016. Methylenebisnicotiflorin: a rare methylene-bridged bisflavonoid glycoside from ripe Pu-er tea. Natural Product Research 30:776−782

doi: 10.1080/14786419.2015.1065491
[30]

Zhu LF, Xu M, Zhu HT, Wang D, Yang SX, et al. 2012. New flavan-3-ol dimer from green tea produced from Camellia taliensis in the Ai-Lao mountains of southwest China. Journal of Agricultural and Food Chemistry 60:12170−12176

doi: 10.1021/jf302726t
[31]

Tian YZ, Liu X, Liu W, Wang WY, Long YH, et al. 2016. A new anti-proliferative acylated flavonol glycoside from Fuzhuan brick-tea. Natural Product Research 30:2637−2641

doi: 10.1080/14786419.2015.1136911
[32]

Yang S, Liu W, Lu S, Tian YZ, Wang WY, et al. 2016. A novel multifunctional compound camellikaempferoside B decreases Aβ production, interferes with Aβ aggregation, and prohibits Aβ-mediated neurotoxicity and neuroinflammation. ACS Chemical Neuroscience 7:505−518

doi: 10.1021/acschemneuro.6b00091
[33]

Lu Y, He Y, Zhu S, Zhong X, Chen D, et al. 2019. New acylglycosides flavones from Fuzhuan brick tea and simulation analysis of their bioactive effects. International Journal of Molecular Sciences 20:494

doi: 10.3390/ijms20030494
[34]

Zhu YF, Chen JJ, Ji XM, Hu X, Ling TJ, et al. 2015. Changes of major tea polyphenols and production of four new B-ring fission metabolites of catechins from post-fermented Jing-Wei Fu brick tea. Food Chemistry 170:110−117

doi: 10.1016/j.foodchem.2014.08.075
[35]

Fu D, Ryan EP, Huang J, Liu Z, Weir TL, et al. 2011. Fermented Camellia sinensis, Fu Zhuan Tea, regulates hyperlipidemia and transcription factors involved in lipid catabolism. Food Research International 44:2999−3005

doi: 10.1016/j.foodres.2011.07.008
[36]

Liu Q, Li J, Chai X, Jiang Y, Tu P. 2013. Chemical constituents from Qianliang tea. Journal of Chinese Pharmaceutical Sciences 2013:427−430

doi: 10.5246/jcps.2013.05.062
[37]

Zhou B, Ma C, Ren X, Xia T, Li X. 2020. LC–MS/MS-based metabolomic analysis of caffeine-degrading fungus Aspergillus sydowii during tea fermentation. Journal of Food Science 85:477−485

doi: 10.1111/1750-3841.15015
[38]

Ling TJ, Wan XC, Ling WW, Zhang ZZ, Xia T, et al. 2010. New triterpenoids and other constituents from a special microbial-fermented tea-Fuzhuan brick tea. Journal of Agricultural and Food Chemistry 58:4945−4950

doi: 10.1021/jf9043524
[39]

Huang YL, Nagai S, Tanaka T, Matsuo Y, Saito Y, et al. 2013. Two new oleanane-type triterpenes isolated from japanese post-fermented tea produced by anaerobic microbial fermentation. Molecules 18:4868−4875

doi: 10.3390/molecules18054868
[40]

Zhu Y, Luo Y, Wang P, Zhao M, Li L, et al. 2016. Simultaneous determination of free amino acids in Pu-erh tea and their changes during fermentation. Food Chemistry 194:643−649

doi: 10.1016/j.foodchem.2015.08.054
[41]

Li Q, Li Y, Luo Y, Zhang Y, Chen Y, et al. 2019. Shifts in diversity and function of the bacterial community during the manufacture of Fu brick tea. Food Microbiology 80:70−76

doi: 10.1016/j.fm.2019.01.001
[42]

Wu YY, Ding L, Xia HL, Tu YY. 2010. Analysis of the major chemical compositions in Fuzhuan brick-tea and its effect on activities of pancreatic enzymes in vitro. African Journal of Biotechnology 9:6748−6754
[43]

Chen G, Peng Y, Xie M, Xu W, Chen C, et al. 2023. A critical review of Fuzhuan brick tea: processing, chemical constituents, health benefits and potential risk. Critical Reviews in Food Science and Nutrition 63:5447−5464

doi: 10.1080/10408398.2021.2020718
[44]

Chen G, Xie M, Wan P, Chen D, Ye H, et al. 2018. Digestion under saliva, simulated gastric and small intestinal conditions and fermentation in vitro by human intestinal microbiota of polysaccharides from Fuzhuan brick tea. Food Chemistry 244:331−339

doi: 10.1016/j.foodchem.2017.10.074
[45]

Hwang LS, Lin LC, Chen NT, Liuchang HC, Shiao MS. 2003. Hypolipidemic effect and antiatherogenic potential of Pu-Erh tea. Oriental Foods and Herbs, eds Ho CT, Lin JK, Zheng QY. Vol. 859. US: ACS Publications. pp. 87−103 doi: 10.1021/bk-2003-0859.ch005
[46]

Aaqil M, Peng C, Kamal A, Nawaz T, Zhang F, et al. 2023. Tea harvesting and processing techniques and its effect on phytochemical profile and final quality of black tea: a review. Foods 12:4467

doi: 10.3390/foods12244467
[47]

Jiao Y, Song Y, Yan Z, Wu Z, Yu Z, et al. 2023. The new insight into the effects of different fixing technology on flavor and bioactivities of orange dark tea. Molecules 28:1079

doi: 10.3390/molecules28031079
[48]

Lv H. 2022. Design and experimental study of steam-heat-air coupling drum tea fixation machine. Thesis. Tai'an: Shandong Agricultural University, China doi: 10.27277/d.cnki.gsdnu.2022.000549
[49]

Moreira J, Aryal J, Guidry L, Adhikari A, Chen Y, et al. 2024. Tea quality: an overview of the analytical methods and sensory analyses used in the most recent studies. Foods 13:3580

doi: 10.3390/foods13223580
[50]

Yang W, Chen R, Sun L, Li Q, Lai X, et al. 2025. Effects of pile-fermentation duration on the taste quality of single-cultivar large-leaf dark tea: insights from metabolomics and microbiomics. Foods 14:670

doi: 10.3390/foods14040670
[51]

Chen QY, Liu ML, Li RY, Jiang B, Liu KY, et al. 2024. Changes in lipids and medium- and long-chain fatty acids during the spontaneous fermentation of ripened pu-erh tea. Current Research in Food Science 9:100831

doi: 10.1016/j.crfs.2024.100831
[52]

Wang T, An J, Bo N, Li R, Chen Q, et al. 2024. Changes and metabolic mechanisms of organic acids in the fermentation of pu-erh tea. LWT 203:116304

doi: 10.1016/j.lwt.2024.116304
[53]

Cheng L, Yang Q, Peng L, Xu L, Chen J, et al. 2024. Exploring core functional fungi driving the metabolic conversion in the industrial pile fermentation of Qingzhuan tea. Food Research International 178:113979

doi: 10.1016/j.foodres.2024.113979
[54]

Yan K, Abbas M, Meng L, Cai H, Peng Z, et al. 2021. Analysis of the fungal diversity and community structure in Sichuan dark tea during pile-fermentation. Frontiers in Microbiology 12:706714

doi: 10.3389/fmicb.2021.706714
[55]

Zhu J, Niu Y, Xiao Z. 2022. Aromatic profiles and enantiomeric distributions of chiral volatile compounds in Pu-erh tea. Journal of Agricultural and Food Chemistry 70:8395−8408

doi: 10.1021/acs.jafc.2c03264
[56]

Li Q, Chai S, Li Y, Huang J, Luo Y, et al. 2018. Biochemical components associated with microbial community shift during the pile-fermentation of primary dark tea. Frontiers in Microbiology 9:1509

doi: 10.3389/fmicb.2018.01509
[57]

Xu J, Wei Y, Li F, Weng X, Wei X. 2022. Regulation of fungal community and the quality formation and safety control of Pu-erh tea. Comprehensive Reviews in Food Science and Food Safety 21:4546−4572

doi: 10.1111/1541-4337.13051
[58]

Wang C, Li J, Wu X, Zhang Y, He Z, et al. 2022. Pu-erh tea unique aroma: volatile components, evaluation methods and metabolic mechanism of key odor-active compounds. Trends in Food Science & Technology 124:25−37

doi: 10.1016/j.tifs.2022.03.031
[59]

Luo Q, Luo L, Zhao J, Wang T, Luo H. 2024. Biological potential and mechanisms of Tea's bioactive compounds: an updated review. Journal of Advanced Research 65:345−363

doi: 10.1016/j.jare.2023.12.004
[60]

Ma C, Ma B, Zhou B, Xu L, Hu Z, et al. 2024. Pile-fermentation mechanism of ripened Pu-erh tea: omics approach, chemical variation and microbial effect. Trends in Food Science & Technology 146:104379

doi: 10.1016/j.tifs.2024.104379
[61]

Luo Y, Zhao Z, Chen H, Pan X, Li R, et al. 2022. Dynamic analysis of physicochemical properties and polysaccharide composition during the pile-fermentation of post-fermented tea. Foods 11:3376

doi: 10.3390/foods11213376
[62]

Wen S, Bai S, An R, Peng Z, Chen H, et al. 2024. Key metabolites influencing astringency and bitterness in Yinghong 9 large-leaf dark tea before and after pile-fermentation. Journal of Agricultural and Food Chemistry 72:27378−27388

doi: 10.1021/acs.jafc.4c07851
[63]

Yang X, Lu B, Mao Q, Yang Q, Liu Z. 2014. Research progress of pile-fermentation of dark tea. Guangdong Agricultural Sciences 41:95−99

doi: 10.16768/j.issn.1004-874x.2014.14.005
[64]

Liu M, Li X, Li Y, Zou Y. 2024. Insights into the airborne microorganisms in a Sichuan south-road dark tea pile fermentation plant during production. Frontiers in Microbiology 15:1439133

doi: 10.3389/fmicb.2024.1439133
[65]

Chen S, Zhang M, Luo S, Ning M, Chen Y, et al. 2025. Multi-omics analysis reveals the sensory quality and fungal communities of Tibetan teas produced by wet- and dry-piling fermentation. Food Research International 201:115690

doi: 10.1016/j.foodres.2025.115690
[66]

Liu SH, Huang FF, Li J, Huang JA, Liu ZH, et al. 2024. The atlas of dark tea: mapping complexities of their microbiome. Trends in Food Science & Technology 154:104780

doi: 10.1016/j.tifs.2024.104780
[67]

Li Y, Hao J, Zhou J, He C, Yu Z, et al. 2022. Pile-fermentation of dark tea: conditions optimization and quality formation mechanism. LWT 166:113753

doi: 10.1016/j.lwt.2022.113753
[68]

Yao Y, Wu M, Huang Y, Li C, Pan X, et al. 2017. Appropriately raising fermentation temperature beneficial to the increase of antioxidant activity and gallic acid content in Eurotium cristatum-fermented loose tea. LWT 82:248−254

doi: 10.1016/j.lwt.2017.04.032
[69]

Wang Z, Su D, Ren H, Lv Q, Ren L, et al. 2022. Effect of different drying methods after fermentation on the aroma of Pu-erh tea (ripe tea). LWT 171:114129

doi: 10.1016/j.lwt.2022.114129
[70]

Zhu W, Wang W, Xu W, Wu S, Chen W, et al. 2022. Influence of thermophilic microorganism on non-volatile metabolites during high-temperature pile-fermentation of Chinese dark tea based on metabolomic analysis. Food Science and Biotechnology 31:827−841

doi: 10.1007/s10068-022-01098-9
[71]

Zhou X, Tian D, Zhou H, Dong R, Ma C, et al. 2024. Effects of different fermentation methods on flavor quality of Liupao tea using GC-Q-TOF-MS and electronic nose analyses. Foods 13:2595

doi: 10.3390/foods13162595
[72]

Feng X, Deng H, Huang L, Teng J, Wei B, et al. 2024. Degradation of cell wall polysaccharides during traditional and tank fermentation of Chinese Liupao tea. Journal of Agricultural and Food Chemistry 72:17692

doi: 10.1021/acs.jafc.4c06264
[73]

Hu Y, Chen W, Gouda M, Yao H, Zuo X, et al. 2024. Fungal fermentation of Fuzhuan brick tea: a comprehensive evaluation of sensory properties using chemometrics, visible near-infrared spectroscopy, and electronic nose. Food Research International 186:114401

doi: 10.1016/j.foodres.2024.114401
[74]

Wang H, Teng J, Huang L, Wei B, Xia N. 2023. Determination of the variations in the metabolic profile and sensory quality of Liupao tea during fermentation through UHPLC-HR-MS metabolomics. Food Chemistry 404:134773

doi: 10.1016/j.foodchem.2022.134773
[75]

Li MY, Xiao Y, Zhong K, Gao H. 2022. Study on taste characteristics and microbial communities in Pingwu Fuzhuan brick tea and the correlation between microbiota composition and chemical metabolites. Journal of Food Science and Technology 59:34−45

doi: 10.1007/s13197-021-04976-y
[76]

Li Q, Jin Y, Jiang R, Xu Y, Zhang Y, et al. 2021. Dynamic changes in the metabolite profile and taste characteristics of Fu brick tea during the manufacturing process. Food Chemistry 344:128576

doi: 10.1016/j.foodchem.2020.128576
[77]

Li J, Wu J, Xu N, Yu Y, Brake J, et al. 2022. Dynamic evolution and correlation between microorganisms and metabolites during manufacturing process and storage of Pu-erh tea. LWT 158:113128

doi: 10.1016/j.lwt.2022.113128
[78]

Wang Z, Zheng C, Ma C, Ma B, Wang J, et al. 2021. Comparative analysis of chemical constituents and antioxidant activity in tea-leaves microbial fermentation of seven tea-derived fungi from ripened Pu-erh tea. LWT 142:111006

doi: 10.1016/j.lwt.2021.111006
[79]

Wang Q, Gong J, Chisti Y, Sirisansaneeyakul S. 2015. Fungal isolates from a Pu-erh type tea fermentation and their ability to convert tea polyphenols to theabrownins. Journal of Food Science 80:M809−M817

doi: 10.1111/1750-3841.12831
[80]

Du Y, Yang W, Yang C, Yang X. 2022. A comprehensive review on microbiome, aromas and flavors, chemical composition, nutrition and future prospects of Fuzhuan brick tea. Trends in Food Science & Technology 119:452−66

doi: 10.1016/j.tifs.2021.12.024
[81]

Zhao P, Park NH, Alam MB, Lee SH. 2022. Fuzhuan brick tea boosts melanogenesis and prevents hair graying through reduction of oxidative stress via NRF2-HO-1 signaling. Antioxidants 11:599

doi: 10.3390/antiox11030599
[82]

Zhang Q, Zhang J. 2024. Overview of Eurotium cristatum and its fermentation application in tea. Future Integrative Medicine 3:11−20

doi: 10.14218/fim.2023.00077
[83]

Wang J, Zhang J, Chen Y, Yu L, Teng J, et al. 2021. The relationship between microbial dynamics and dominant chemical components during Liupao tea processing. Food Bioscience 43:101315

doi: 10.1016/j.fbio.2021.101315
[84]

Wu S, Wang W, Zhu W, Chen W, Xu W, et al. 2022. Microbial community succession in the fermentation of Qingzhuan tea at various temperatures and their correlations with the quality formation. International Journal of Food Microbiology 382:109937

doi: 10.1016/j.ijfoodmicro.2022.109937
[85]

Zhu H, Zhou X, Shen C, Ao Z, Cao X, et al. 2024. Bacillus licheniformis-based intensive fermentation of Tibetan tea improved its bioactive compounds and reinforced the intestinal barrier in mice. Frontiers in Microbiology 15:1376757

doi: 10.3389/fmicb.2024.1376757
[86]

Cheng L, Peng L, Li X, Xu L, Chen J, et al. 2024. Co-occurrence network and functional profiling of the bacterial community in the industrial pile fermentation of Qingzhuan tea: understanding core functional bacteria. Food Chemistry 454:139658

doi: 10.1016/j.foodchem.2024.139658
[87]

Tian D, Huang G, Ren L, Li Y, Yu J, et al. 2024. Effects of Monascus purpureus on ripe Pu-erh tea in different fermentation methods and identification of characteristic volatile compounds. Food Chemistry 440:138249

doi: 10.1016/j.foodchem.2023.138249
[88]

Chen Q, Zhang M, Chen M, Li M, Zhang H, et al. 2021. Influence of Eurotium cristatum and Aspergillus niger individual and collaborative inoculation on volatile profile in liquid-state fermentation of instant dark teas. Food Chemistry 350:129234

doi: 10.1016/j.foodchem.2021.129234
[89]

Xu Q, Sun M, Ning J, Fang S, Ye Z, et al. 2019. The core role of Bacillus subtilis and Aspergillus fumigatus in pile-fermentation processing of Qingzhuan brick tea. Indian Journal of Microbiology 59:288−294

doi: 10.1007/s12088-019-00802-4
[90]

Cheng L, Peng L, Xu L, Yu X, Zhu Y, et al. 2025. Metabolic function and quality contribution of tea-derived microbes, and their safety risk in dark tea manufacture. Food Chemistry 464:141818

doi: 10.1016/j.foodchem.2024.141818
[91]

Yu Z, Deng H, Qu H, Zhang B, Lei G, et al. 2022. Penicillium simplicissimum possessing high potential to develop decaffeinated Qingzhuan tea. LWT 165:113606

doi: 10.1016/j.lwt.2022.113606
[92]

Wang Y, Zhang M, Zhang Z, Lu H, Gao X, et al. 2017. High-theabrownins instant dark tea product by Aspergillus niger via submerged fermentation: α-glucosidase and pancreatic lipase inhibition and antioxidant activity. Journal of the Science of Food and Agriculture 97:5100−5106

doi: 10.1002/jsfa.8387
[93]

Ma W, Zhu Y, Shi J, Wang J, Wang M, et al. 2021. Insight into the volatile profiles of four types of dark teas obtained from the same dark raw tea material. Food Chemistry 346:128906

doi: 10.1016/j.foodchem.2020.128906
[94]

Jiao Y, Tang H, Yan Z, Wu Z, Zhang D, et al. 2024. Effect of different drying methods on quality of orange dark tea. Journal of Food Measurement and Characterization 18:3244−3254

doi: 10.1007/s11694-024-02399-9
[95]

Liu P, Feng L, Chen J, Wang S, Wang X, et al. 2024. Unlocking the secrets of Qingzhuan tea: a comprehensive overview of processing, flavor characteristics, and health benefits. Trends in Food Science & Technology 147:104450

doi: 10.1016/j.tifs.2024.104450
[96]

Pang Y, Wei Y, Wei K, Huang R, Peng L, et al. 2024. A rising star in dark teas: the processing, microorganisms, chemicals, bioactivities, and safety of Qingzhuan tea. ACS Food Science & Technology 4:796−812

doi: 10.1021/acsfoodscitech.3c00630