[1]

de Almeida Teixeira GH, Berlingieri Durigan MF, Durigan JF. 2011. Jaboticaba (Myrciaria cauliflora (Mart.) O. Berg. [Myrtaceae]). In Postharvest Biology and Technology of Tropical and Subtropical Fruits, ed. Yahia EM. UK: Woodhead Publishing. pp. 246−274,275e doi: 10.1533/9780857092885.246

[2]

Castangia I, Manca ML, Allaw M, Hellström J, Granato D, et al. 2021. Jabuticaba (Myrciaria jaboticaba) peel as a sustainable source of anthocyanins and ellagitannins delivered by phospholipid vesicles for alleviating oxidative stress in human keratinocytes. Molecules 26:6697

doi: 10.3390/molecules26216697
[3]

Inada KOP, Oliveira AA, Revorêdo TB, Martins ABN, Lacerda ECQ, et al. 2015. Screening of the chemical composition and occurring antioxidants in jabuticaba (Myrciaria jaboticaba) and Jussara (Euterpe edulis) fruits and their fractions. Journal of Functional Foods 17:422−433

doi: 10.1016/j.jff.2015.06.002
[4]

Pereira LD, Barbosa JM, Ribeiro da Silva AJ, Ferri PH, Santos SC. 2017. Polyphenol and ellagitannin constituents of jabuticaba (Myrciaria cauliflora) and chemical variability at different stages of fruit development. Journal of Agricultural and Food Chemistry 65:1209−1219

doi: 10.1021/acs.jafc.6b02929
[5]

Plaza M, Batista ÂG, Cazarin CBB, Sandahl M, Turner C, et al. 2016. Characterization of antioxidant polyphenols from Myrciaria jaboticaba peel and their effects on glucose metabolism and antioxidant status: a pilot clinical study. Food Chemistry 211:185−197

doi: 10.1016/j.foodchem.2016.04.142
[6]

Wu SB, Dastmalchi K, Long C, Kennelly EJ. 2012. Metabolite profiling of jaboticaba (Myrciaria cauliflora) and other dark-colored fruit juices. Journal of Agricultural and Food Chemistry 60:7513−7525

doi: 10.1021/jf301888y
[7]

Macedo EHBC, Santos GC Jr, Santana MN, Jesus EFO, de Araújo UB, et al. 2021. Unveiling the physicochemical properties and chemical profile of artisanal jabuticaba wines by bromatological and NMR-based metabolomics approaches. LWT 146:111371

doi: 10.1016/j.lwt.2021.111371
[8]

Skates E, Overall J, DeZego K, Wilson M, Esposito D, et al. 2018. Berries containing anthocyanins with enhanced methylation profiles are more effective at ameliorating high fat diet-induced metabolic damage. Food and Chemical Toxicology 111:445−453

doi: 10.1016/j.fct.2017.11.032
[9]

Yang L, Ling W, Du Z, Chen Y, Li D, et al. 2017. Effects of anthocyanins on cardiometabolic health: a systematic review and meta-analysis of randomized controlled trials. Advances in Nutrition 8:684−693

doi: 10.3945/an.116.014852
[10]

Giampieri F, Gasparrini M, Forbes-Hernandez TY, Mazzoni L, Capocasa F, et al. 2018. Overexpression of the anthocyanidin synthase gene in strawberry enhances antioxidant capacity and cytotoxic effects on human hepatic cancer cells. Journal of Agricultural and Food Chemistry 66:581−592

doi: 10.1021/acs.jafc.7b04177
[11]

Giampieri F, Alvarez-Suarez JM, Mazzoni L, Forbes-Hernandez TY, Gasparrini M, et al. 2014. An anthocyanin-rich strawberry extract protects against oxidative stress damage and improves mitochondrial functionality in human dermal fibroblasts exposed to an oxidizing agent. Food & Function 5:1939−1948

doi: 10.1039/C4FO00048J
[12]

Rahman MM, Ichiyanagi T, Komiyama T, Sato S, Konishi T. 2008. Effects of anthocyanins on psychological stress-induced oxidative stress and neurotransmitter status. Journal of Agricultural and Food Chemistry 56:7545−7550

doi: 10.1021/jf800930s
[13]

Ranganath KG, Shivashankara KS, Roy TK, Dinesh MR, Geetha GA, et al. 2018. Profiling of anthocyanins and carotenoids in fruit peel of different colored mango cultivars. Journal of Food Science and Technology 55:4566−4577

doi: 10.1007/s13197-018-3392-7
[14]

Sadowska-Bartosz I, Bartosz GJ. 2024. Antioxidant activity of anthocyanins and anthocyanidins: a critical review. International Journal of Molecular Sciences 25:12001

doi: 10.3390/ijms252212001
[15]

Xu Y, Li H, Shi T, Luo Q, Chen Y, et al. 2025. High-quality genome of black wolfberry (Lycium ruthenicum Murr.) provides insights into the genetics of anthocyanin biosynthesis regulation. Horticulture Research 12:uhae298

doi: 10.1093/hr/uhae298
[16]

Duan X, Wang K, Tang R, Liu J, Cheng K, et al. 2025. Recent advances in biosynthesis and regulation of strawberry anthocyanins. Horticulture Research 12(8):uhaf135

doi: 10.1093/hr/uhaf135
[17]

Zhang L, Song B, Li B, Zhang S, Liu Y, et al. 2024. Genome-wide identification and expression analysis of fifteen gene families involved in anthocyanin synthesis in pear. Horticulturae 10:335

doi: 10.3390/horticulturae10040335
[18]

Jaakola L. 2013. New insights into the regulation of anthocyanin biosynthesis in fruits. Trends in Plant Science 18:477−483

doi: 10.1016/j.tplants.2013.06.003
[19]

Koes R, Verweij W, Quattrocchio F. 2005. Flavonoids: a colorful model for the regulation and evolution of biochemical pathways. Trends in Plant Science 10:236−242

doi: 10.1016/j.tplants.2005.03.002
[20]

Winkel-Shirley B. 2001. Flavonoid biosynthesis. A colorful model for genetics, biochemistry, cell biology, and biotechnology. Plant Physiology 126:485−493

doi: 10.1104/pp.126.2.485
[21]

Zhang H, Yang B, Liu J, Guo D, Hou J, et al. 2017. Analysis of structural genes and key transcription factors related to anthocyanin biosynthesis in potato tubers. Scientia Horticulturae 225:310−316

doi: 10.1016/j.scienta.2017.07.018
[22]

Wu Z, Bian R, Zhang Z, Li L, Zhang J. 2025. Transcription factors repressing anthocyanin biosynthesis in horticultural crops. Fruit Research 5:e007

doi: 10.48130/frures-0024-0042
[23]

Guo Y, Zhang H, Shao S, Sun S, Yang D, et al. 2022. Anthocyanin: a review of plant sources, extraction, stability, content determination and modifications. International Journal of Food Science & Technology 57:7573−7591

doi: 10.1111/ijfs.16132
[24]

Bocker R, Silva EK. 2024. Anthocyanin-rich jaboticaba fruit: Natural source of bioactive and coloring ingredients for nutraceutical food applications. Trends in Food Science & Technology 153:104744

doi: 10.1016/j.tifs.2024.104744
[25]

Leite-Legatti AV, Batista ÂG, Dragano NRV, Marques AC, Malta LG, et al. 2012. Jaboticaba peel: antioxidant compounds, antiproliferative and antimutagenic activities. Food Research International 49:596−603

doi: 10.1016/j.foodres.2012.07.044
[26]

de Souza ÁC, Geraldi MV, Marostica MR Jr. 2025. Jaboticaba berry: metabolic benefits, bioactive compounds, and opportunities for commercialization. Trends in Food Science & Technology 157:104894

doi: 10.1016/j.tifs.2025.104894
[27]

Trainin T, Harel-Beja R, Bar-Ya'akov I, Ben-Simhon Z, Yahalomi R, et al. 2021. Fine mapping of the "black" peel color in pomegranate (Punica granatum L.) strongly suggests that a mutation in the anthocyanidin reductase (ANR) gene is responsible for the trait. Frontiers in Plant Science 12:642019

doi: 10.3389/fpls.2021.642019
[28]

Paun N, Botoran OR, Niculescu VC. 2022. Total phenolic, anthocyanins HPLC-DAD-MS determination and antioxidant capacity in black grape skins and blackberries: a comparative study. Applied Sciences 12:936

doi: 10.3390/app12020936
[29]

Condurache Lazăr NN, Croitoru C, Enachi E, Bahrim GE, Stănciuc N, et al. 2021. Eggplant peels as a valuable source of anthocyanins: extraction, thermal stability and biological activities. Plants 10:577

doi: 10.3390/plants10030577
[30]

Wang XY, Zhang C, Zhang M, Ma CC, Yan JP, et al. 2022. Study of anthocyanin composition and fruit color in Osmanthus fragrans 'Zi Yingui'. Plant Science Journal 40(5):677−687

doi: 10.11913/PSJ.2095-0837.2022.50677
[31]

Cao L, Park Y, Lee S, Kim DO. 2021. Extraction, identification, and health benefits of anthocyanins in blackcurrants (Ribes nigrum L.). Applied Sciences 11:1863

doi: 10.3390/app11041863
[32]

He S, Dong W, Chen J, Zhang J, Lin W, et al. 2024. DataColor: unveiling biological data relationships through distinctive color mapping. Horticulture Research 11:uhad273

doi: 10.1093/hr/uhad273
[33]

Li Z, Wang C, Wang S, Wang W, Chen F. 2024. HortDB V1.0: a genomic database of horticultural plants. Horticulture Research 11:uhae224

doi: 10.1093/hr/uhae224
[34]

Barnes JS, Nguyen HP, Shen S, Schug KA. 2009. General method for extraction of blueberry anthocyanins and identification using high performance liquid chromatography–electrospray ionization-ion trap-time of flight-mass spectrometry. Journal of Chromatography A 1216:4728−4735

doi: 10.1016/j.chroma.2009.04.032
[35]

Glauser G, Grund B, Gassner AL, Menin L, Henry H, et al. 2016. Validation of the mass-extraction-window for quantitative methods using liquid chromatography high resolution mass spectrometry. Analytical Chemistry 88:3264−3271

doi: 10.1021/acs.analchem.5b04689
[36]

Lin Z, Deng K, Chen F, Zhao LJH. 2025. Integrated multi-omics reveals anthocyanin biosynthesis control during fruit peel color development in jaboticaba 'Essart'. Horticulturae 11:1515

doi: 10.3390/horticulturae11121515
[37]

Zhao L, Li Z, Jiang S, Xia C, Deng K, et al. 2024. The telomere-to-telomere genome of jaboticaba reveals the genetic basis of fruit color and citric acid content. International Journal of Molecular Sciences 25:11951

doi: 10.3390/ijms252211951
[38]

Zhang J, Zheng Y, Chen F. 2025. Phased telomere-to-telomere super-pangenome: definitive reference genome in plants. Trends in Plant Science 31(3):266−269

doi: 10.1016/j.tplants.2025.11.002
[39]

Chen F, Song Y, Li X, Chen J, Mo L, et al. 2019. Genome sequences of horticultural plants: past, present, and future. Horticulture Research 6:112

doi: 10.1038/s41438-019-0195-6
[40]

Niu F, Cui X, Zhao P, Sun M, Yang B, et al. 2020. WRKY42 transcription factor positively regulates leaf senescence through modulating SA and ROS synthesis in Arabidopsis thaliana. The Plant Journal 104:171−184

doi: 10.1111/tpj.14914
[41]

Borevitz JO, Xia Y, Blount J, Dixon RA, Lamb C. 2000. Activation tagging identifies a conserved MYB regulator of phenylpropanoid biosynthesis. The Plant Cell 12:2383−2394

doi: 10.1105/tpc.12.12.2383
[42]

Zuluaga DL, Gonzali S, Loreti E, Pucciariello C, Degl'Innocenti E, et al. 2008. Arabidopsis thaliana MYB75/PAP1 transcription factor induces anthocyanin production in transgenic tomato plants. Functional Plant Biology 35:606−618

doi: 10.1071/FP08021
[43]

Hellens RP, Allan AC, Friel EN, Bolitho K, Grafton K, et al. 2005. Transient expression vectors for functional genomics, quantification of promoter activity and RNA silencing in plants. Plant Methods 1:13

doi: 10.1186/1746-4811-1-13
[44]

Wang F, Chen J, Tang R, Wang R, Ahmad S, et al. 2023. Research progress on anthocyanin-mediated regulation of 'black' phenotypes of plant organs. Current Issues in Molecular Biology 45:7242−7256

doi: 10.3390/cimb45090458
[45]

Petruskevicius A, Viskelis J, Urbonaviciene D, Viskelis P. 2023. Anthocyanin accumulation in berry fruits and their antimicrobial and antiviral properties: an overview. Horticulturae 9:288

doi: 10.3390/horticulturae9020288
[46]

Ponder A, Hallmann E, Kwolek M, Średnicka-Tober D, Kazimierczak R. 2021. Genetic differentiation in anthocyanin content among berry fruits. Current Issues in Molecular Biology 43:36−51

doi: 10.3390/cimb43010004
[47]

Huo J, Ni Y, Li D, Qiao J, Huang D, et al. 2023. Comprehensive structural analysis of polyphenols and their enzymatic inhibition activities and antioxidant capacity of black mulberry (Morus nigra L.). Food Chemistry 427:136605

doi: 10.1016/j.foodchem.2023.136605
[48]

Enaru B, Drețcanu G, Pop TD, Stǎnilǎ A, Diaconeasa Z. 2021. Anthocyanins: factors affecting their stability and degradation. Antioxidants 10:1967

doi: 10.3390/antiox10121967
[49]

Trouillas P, Sancho-García JC, De Freitas V, Gierschner J, Otyepka M, et al. 2016. Stabilizing and modulating color by copigmentation: insights from theory and experiment. Chemical Reviews 116:4937−4982

doi: 10.1021/acs.chemrev.5b00507
[50]

Passeri V, Koes R, Quattrocchio FM. 2016. New challenges for the design of high value plant products: stabilization of anthocyanins in plant vacuoles. Frontiers in Plant Science 7:153

doi: 10.3389/fpls.2016.00153
[51]

Khoo HE, Azlan A, Tang ST, Lim SM. 2017. Anthocyanidins and anthocyanins: Colored pigments as food, pharmaceutical ingredients, and the potential health benefits. Food & Nutrition Research 61:1361779

doi: 10.1080/16546628.2017.1361779
[52]

Sigurdson GT, Giusti MM. 2014. Bathochromic and hyperchromic effects of aluminum salt complexation by anthocyanins from edible sources for blue color development. Journal of Agricultural and Food Chemistry 62:6955−6965

doi: 10.1021/jf405145r
[53]

Montefiori M, Comeskey DJ, Wohlers M, McGhie TK. 2009. Characterization and quantification of anthocyanins in red kiwifruit (Actinidia spp.). Journal of Agricultural and Food Chemistry 57:6856−6861

doi: 10.1021/jf900800z
[54]

Freyre R, Uzdevenes C, Gu L, Quesenberry KH. 2015. Genetics and anthocyanin analysis of flower color in Mexican Petunia. Journal of the American Society for Horticultural Science 140:45−49

doi: 10.21273/jashs.140.1.45
[55]

Nutricati E, Sabella E, Negro C, Min Allah S, Luvisi A, et al. 2025. Anthocyanins and anthocyanin biosynthesis gene expression in Passiflora flower corona filaments. Plants 14:1050

doi: 10.3390/plants14071050
[56]

Wang J, Zhao Y, Sun B, Yang Y, Wang S, et al. 2024. The structure of anthocyanins and the copigmentation by common micromolecular copigments: a review. Food Research International 176:113837

doi: 10.1016/j.foodres.2023.113837
[57]

Li Y, Wang J, Lu C, Wang Z, Deng C, et al. 2024. Flavonoid extracts from chrysanthemum with appropriate anthocyanins turn blue when exposed to iron ions. Horticultural Plant Journal 10:837−852

doi: 10.1016/j.hpj.2023.10.002
[58]

He F, Mu L, Yan GL, Liang NN, Pan QH, et al. 2010. Biosynthesis of anthocyanins and their regulation in colored grapes. Molecules 15:9057−9091

doi: 10.3390/molecules15129057
[59]

Ramsay NA, Glover BJ. 2005. MYB–bHLH–WD40 protein complex and the evolution of cellular diversity. Trends in Plant Science 10:63−70

doi: 10.1016/j.tplants.2004.12.011
[60]

Bulanov AN, Andreeva EA, Tsvetkova NV, Zykin PA. 2025. Regulation of flavonoid biosynthesis by the MYB-bHLH-WDR (MBW) complex in plants and its specific features in cereals. International Journal of Molecular Sciences 26:734

doi: 10.3390/ijms26020734
[61]

Pireyre M, Burow M. 2015. Regulation of MYB and bHLH transcription factors: a glance at the protein level. Molecular Plant 8:378−388

doi: 10.1016/j.molp.2014.11.022
[62]

Cui D, Zhao S, Xu H, Allan AC, Zhang X, et al. 2021. The interaction of MYB, bHLH and WD40 transcription factors in red pear (Pyrus pyrifolia) peel. Plant Molecular Biology 106:407−417

doi: 10.1007/s11103-021-01160-w
[63]

Li S. 2014. Transcriptional control of flavonoid biosynthesis: fine-tuning of the MYB-bHLH-WD40 (MBW) complex. Plant Signaling & Behavior 9:e27522

doi: 10.4161/psb.27522
[64]

Zhang L, Wang Y, Sun M, Wang J, Kawabata S, et al. 2014. BrMYB4, a suppressor of genes for phenylpropanoid and anthocyanin biosynthesis, is down-regulated by UV-B but not by pigment-inducing sunlight in turnip cv. Tsuda. Plant & Cell Physiology 55:2092−2101

doi: 10.1093/pcp/pcu137
[65]

Wang XC, Wu J, Guan ML, Zhao CH, Geng P, et al. 2020. Arabidopsis MYB4 plays dual roles in flavonoid biosynthesis. The Plant Journal 101:637−652

doi: 10.1111/tpj.14570