doi:10.48130/mpb-0026-0017

[1]

Editorial Committee of Flora Reipublicae Popularis Sinicae, Chinese Academy of Sciences. 1989. Flora Reipublicae Popularis Sinicae. Vol. 64(2). Beijing: Science Press. pp. 42 www.iplant.cn/foc

[2]

Bao S, Bi Y, Shan F, Eni R, Zhong X, et al. 2026. A comprehensive review on botany, phytochemistry, pharmacology, and resource protection of genus Arnebia Forssk. Phytochemistry Reviews 25(2):1813−1856

doi: 10.1007/s11101-025-10182-x

[3]

Chinese Pharmacopoeia Commission. 2025. 中华人民共和国药典: 一部 [Pharmacopoeia of the People's Republic of China: volume I]. 北京 [Beijing]: 中国医药科技出版社 [China Medical Science and Technology Press] pp. 128 (in Chinese) https://2025.chp.org.cn/#/main

[4]

Yusupu A, Zhou LN, Guo YQ, Abudukelimu M, Tuerhong M. 2022. 黄花软紫草的多糖成分及其抗病毒活性研究 [Studies on polysaccharide from Arnebia guttata bunge and its antiviral activity]. 南开大学学报(自然科学版) [Journal of Nankai University (Natural Science)] 55:86−91 (in Chinese)

[5]

Yuan MJ, Zhu XY, Shi HS, Zhao Z, Fan WJ, et al. 2025. 紫草素治疗糖尿病足溃疡的疗效及基于网络药理学的作用机制 [Therapeutic effects and mechanisms of Shikonin on diabetic foot ulcers in mice based on network pharmacology]. 世界中医药 [World Chinese Medicine] 20:2239−2248 (in Chinese)

doi: 10.3969/j.issn.1673-7202.2025.13.003

[6]

Xuan XF, Li GC. 2025. 黄连紫草膏结合高压氧对老年糖尿病足溃疡患者炎症因子与创面修复效果的影响 [Effect of Huanglian Zicao ointment combined with hyperbaric oxygen on inflammatory factors and wound repair in elderly patients with diabetic foot ulcers]. 中国老年学杂志 [Chinese Journal of Gerontology] 45(19):4717−4721 (in Chinese)

doi: 10.3969/j.issn.1005-9202.2025.19.030

[7]

Bao M, Liu MY, Pan XZ, Lei N, Chen Q, et al. 2025. β-羟基异戊酰紫草素对卵巢癌裸鼠移植瘤生长的影响及作用机制 [Effects and mechanism of β-hydroxyisovalerylshikonin on the growth of ovarian cancer xenografts in nude mice]. 中国老年学杂志 [Chinese Journal of Gerontology] 45(17):4257−4261 (in Chinese)

doi: 10.3969/j.issn.1005-9202.2025.17.040

[8]

Cheng BX, Yan J, Han YY, Qu BH, Guang ZX, et al. 2025. 紫草素治疗结直肠癌作用机制研究进展 [Research progress on the mechanism of action of shikonin in the treatment of colorectal cancer]. 中华中医药学刊 [Chinese Archives of Traditional Chinese Medicine] 00:1−15 (in press) (in Chinese)

[9]

Yan YM, Zou LJ, Li MH, Sun ZC, Zhang M, et al. 2023. Three new compounds with their anti-glioma effects from the roots of Arnebia guttata Bunge. Natural Product Research 37(17):2817−2823

doi: 10.1080/14786419.2022.2136658

[10]

National Forestry and Grassland Administration, Ministry of Agriculture and Rural Affairs. 2021. 国家重点保护野生植物名录 [National List of Key Protected Wild Plants]. 中国政府网 [Chinese Government Website] (in Chinese) www.gov.cn/zhengce/zhengceku/2021-09/09/content_5636409.htm

[11]

Ding WH, Wang DD, Wang YJ, Li J, Xu HY. 2023. 新疆3种软紫草资源现状调查与分析 [Investigation and analysis of three species of Arnebia resource situation in Xinjiang]. 时珍国医国药 [Journal of Li-shizhen Traditional Chinese Medicine] 34:162−165 (in Chinese)

doi: 10.3969/j.issn.1008-0805.2023.01.44

[12]

Dai SY, Liu J, Yuan SN, Lian CJ, Qiao F, et al. 2024. 基于2015年和2022年国家药品抽检探讨紫草的质量现状 [Discussion on the quality status of Arnebiae Radix based on the national drug sampling and testing in 2015 and 2022]. 药物分析杂志 [Chinese Journal of Pharmaceutical Analysis] 44:740−749 (in Chinese)

doi: 10.16155/j.0254-1793.2024.05.01

[13]

Liu JW, Li XZ, Yao MJ. 2021. 植物根际微生物群落构建的研究进展 [Research progress on assembly of plant rhizosphere microbial community]. 微生物学报 [Acta Microbiologica Sinica] 61(2):231−246 (in Chinese)

doi: 10.13343/j.cnki.wsxb.20200154

[14]

Shi M, Hao S, Wang Y, Zhang S, Cui G, et al. 2024. Plant growth-promoting fungi improve tobacco yield and chemical components by reassembling rhizosphere fungal microbiome and recruiting probiotic taxa. Environmental Microbiome 19(1):83

doi: 10.1186/s40793-024-00629-7

[15]

Mendes R, Garbeva P, Raaijmakers JM. 2013. The rhizosphere microbiome: significance of plant beneficial, plant pathogenic, and human pathogenic microorganisms. FEMS Microbiology Reviews 37(5):634−663

doi: 10.1111/1574-6976.12028

[16]

Lugtenberg B, Kamilova F. 2009. Plant-growth-promoting rhizobacteria. Annual Review of Microbiology 63(1):541−556

doi: 10.1146/annurev.micro.62.081307.162918

[17]

Luo SY, Tian CJ, Zhang JF, Ji L, Yao ZM, et al. 2022. 根际微生物对药材道地性的影响及应用前景展望 [Influence of inter-root microorganisms on the authenticity of medicinal herbs and prospects for their application]. 时珍国医国药 [Lishizhen Medicine and Materia Medica Research] 33:948−950 (in Chinese)

doi: 10.3969/j.issn.1008-0805.2022.04.51

[18]

Zhang SM, Huang YM, Ni YX, Zhong QQ. 2018. 宁南山区人工林草对土壤真菌群落的影响 [Effects of artificial forest and grass on soil fungal community at southern Ningxia mountain]. 中国环境科学 [China Environmental Science] 38(4):1449−1458 (in Chinese)

doi: 10.19674/j.cnki.issn1000-6923.2018.0175

[19]

Xu JM, Zhou YH, Gao XM, Yang CX. 2022. 盐胁迫下丛枝菌根真菌对蒲公英矿质元素吸收的影响 [Effects of arbuscular mycorrhizal fungi on mineral element absorption in dandelion under salt stress]. 山西农业科学 [Journal of Shanxi Agricultural Sciences] 50(2):206−212 (in Chinese)

doi: 10.3969/j.issn.1002-2481.2022.02.11

[20]

Li N. 2025. 微生物菌肥在农业生产中的应用 [Application of microbial fertilizer in agricultural production]. 种子科技 [Seed Science and Technology] 43:213−215 (in Chinese)

doi: 10.19904/j.cnki.cn14-1160/s.2025.19.069

[21]

Wu QM, Zhang JM, Li YY, Zhang Y. 2021. 有益微生物菌肥对农作物的作用机制研究进展 [Research progress on the mechanism of action of beneficial microbial fertilizers on crops]. 生物技术通报 [Biotechnology Bulletin] 37(5):221−230 (in Chinese)

doi: 10.13560/j.cnki.biotech.bull.1985.2020-0846

[22]

Zhang ZJ, Qiu YJ, Zhao YQ, Ye Y, Wang G, et al. 2023. 新疆紫草根际环境与药材药效成分含量相关性研究 [Correlation study between rhizosphere environment and content of medicinal components in Lithospermum erythrorhizon, Xinjiang]. 中国中药杂志 [China Journal of Chinese Materia Medica] 48(22):6030−6038 (in Chinese)

doi: 10.19540/j.cnki.cjcmm.20230725.101

[23]

Li Y, Zhao W, Zuo X, Ni B, You J, et al. 2021. After habitat changed, evaluation for genetic diversity and secondary metabolic changes of cultivated and wild populations of medicinal plants. Research Square Preprint

doi: 10.21203/rs.3.rs-864658/v1

[24]

Tian CF, Hu QN, Zhan ZL, Lan XY, Li X, et al. 2024. 基于传统品质评价的野生与栽培赤芍比较 [Comparison of wild and cultivated Paeoniae radix rubra based on traditional quality evaluation]. 中国实验方剂学杂志 [Chinese Journal of Experimental Traditional Medical Formulae] 30(14):165−174 (in Chinese)

doi: 10.13422/j.cnki.syfjx.20240467

[25]

Li Y, Zhao W, Zuo X, You J, Xing M, et al. 2025. Habitat changes due to cultivation alter the genetic diversity and secondary metabolic changes of medicinal plants. Scientific Reports 15(1):36185

doi: 10.1038/s41598-025-20010-z

[26]

Shang YJ, Geng B, Chen X, Wang Q, Zheng GH, et al. 2026. 基于传统品质评价的野生与栽培栀子品质比较 [Comparison of wild and cultivated Gardeniae Fructus based on traditional quality evaluation]. 中国实验方剂学杂志 [Chinese Journal of Experimental Traditional Medical Formulae] 32(5):225−234 (in Chinese)

doi: 10.13422/j.cnki.syfjx.20251665

[27]

Ding WH, Yan XH, Ge L, Xu HY. 2019. HPLC法测定新疆软紫草属4种植物根中萘醌类成分 [Determination of naphthoquinones in roots of four species of Xinjiang soft comfrey by HPLC method]. 中成药 [Chinese Traditional Patent Medicine] 41:936−939 (in Chinese)

doi: 10.3969/j.issn.1001-1528.2019.04.047

[28]

Eller F, Guo X, Ye S, Mozdzer TJ, Brix H. 2020. Suitability of wild Phragmites australis as bio-resource: tissue quality and morphology of populations from three continents. Resources 9(12):143

doi: 10.3390/resources9120143

[29]

Eds.

Bao SD. 2010. 土壤农化分析 [Soil agrochemical analysis]. 中国农业出版社 [China Agricultural Press] pp. 14−114 (in Chinese)

[30]

Xing Y, Wang X. 2024. Precise application of water and fertilizer to crops: challenges and opportunities. Frontiers in Plant Science 15:1444560

doi: 10.3389/fpls.2024.1444560

[31]

Sun W, Li Q, Qiao B, Jia K, Li C, et al. 2024. Advances in plant–soil feedback driven by root exudates in forest ecosystems. Forests 15(3):515

doi: 10.3390/f15030515

[32]

Jalal A, da Silva Oliveira CE, Gato IMB, de Almeida Moreira V, de Lima BH, et al. 2025. Interaction of mineral nutrients and plant growth-promoting microbes for biofortification of different cropping systems. Journal of Plant Growth Regulation 44(12):6785−6801

doi: 10.1007/s00344-024-11380-1

[33]

Ng CWW, So PS, Coo JL, Lou SY, Wong JTF. 2023. Interactions between nutrient types and soil hydrological properties on yield and quality of Pinellia ternata, a medicinal plant. Industrial Crops and Products 195(2):116423

doi: 10.1016/j.indcrop.2023.116423

[34]

Liu S, Xiong L, Fang W, Wang K, Cui X, et al. 2025. Effects of nitrogen, phosphorous, and potassium fertilization on rapeseed yield under freeze stress. Crop and Environment 4(3):143−153

doi: 10.1016/j.crope.2025.04.002

[35]

Xia F, Liu YN, Lu X, Fu H, Chen Y, et al. 2025. 不同基肥及施用量对设施大棚土壤养分和微生物的影响 [Effects of different base fertilizers and application rates on soil nutrients and microorganisms in facility greenhouses]. 热带作物学报 [Chinese Journal of Tropical Crops] 46(8):2019−2029 (in Chinese)

doi: 10.3969/j.issn.1000-2561.2025.08.024

[36]

Lin H, Wu T, Li YJ. 2026. 油橄榄根际细菌群落与叶片及土壤营养状况的关联性 [Correlation between rhizosphere bacterial community and leaf & soil nutrient status of Europaea oleopaea]. 经济林研究 [Non-wood Forest Research] 4(1):33−45 (in Chinese)

doi: 10.14067/j.cnki.1003-8981.2026.01.004

[37]

Li RT, Yao HK, Liu YF, Yang SD. 2016. Effect of low phosphorus stress on root growth and soil bacterial diversity in rhizosphere of tomatoes. Acta Horticulturae Sinica 43:473−484

doi: 10.16420/j.issn.0513-353x.2015-0919

[38]

Wang C, Tai H, Chen Y, Zhai Z, Zhang L, et al. 2025. Soil microbiota modulates root transcriptome with divergent effect on maize growth under low and high phosphorus inputs. Plant, Cell & Environment 48(3):2132−2144

doi: 10.1111/pce.15281

[39]

Chen F, Wang XS, Gan GY, Li YL, Zou JL, et al. 2021. 长期施用磷肥对稻-油轮作土壤磷组分及微生物多样性的影响 [Effects of long-term application of phosphorus fertilizer on soil phosphorus fractions and microbial diversity in rice-rapeseed rotation]. 华中农业大学学报 [Journal of Huazhong Agricultural University] 40:168−178 (in Chinese)

doi: 10.13300/j.cnki.hnlkxb.2021.01.021

[40]

Li YZ, Tan ZY, Liu J, Yang J, Hu Q, et al. 2026. 不同微生物菌剂对植烟土壤微生物群落结构及功能的影响 [Effects of different microbial agents on the microbial community structure and functions of tobacco-planting soil]. 中国土壤与肥料 [Soil and Fertilizer Sciences in China] 123(4):82−95 (in Chinese)

doi: 10.11838/sfsc.1673-6257.25418

[41]

Chao FC, Zhang FS, Yang YH, Li CJ. 2003. 影响烟草根系发育的几个因素探讨 [Effect of factors on the growth of tobacco root]. 中国烟草科学 [Chinese Tobacco Science] 24(2):6−9 (in Chinese)

doi: 10.13496/j.issn.1007-5119.2003.02.002

[42]

Zhang SW, Chen KK. 2025. Research progress on effect of low phosphorus stress on tobacco growth and physiological characteristics. Modern Agricultural Science and Technology (22):65−71

doi: 10.3969/j.issn.1007-5739.2025.22.018

[43]

Zeng L, Yang G, Fang H, Chen J, Ma X. 2026. 土壤有效磷调控玉米根际碳动态的植物-微生物互作机制 [Plant–microbe interaction mechanism of soil available phosphorus in regulating carbon dynamics in maize rhizosphere]. 浙江农林大学学报 [Journal of Zhejiang A&F University] 43(1):133−141 (in Chinese)

doi: 10.11833/j.issn.2095-0756.20250112

[44]

Ding N, Lin H, Zhang XH, He Y, Yu G. 2022. 植物根系分泌物与根际微生物交互作用机制研究进展 [Interaction mechanism between root secretion and rhizosphere microorganisms: a review]. 土壤通报 [Chinese Journal of Soil Science] 53(5):1212−1219 (in Chinese)

doi: 10.19336/j.cnki.trtb.2022010201

[45]

Tao DX, Gao YZ. 2023. 土壤解磷微生物促进植物磷素吸收策略研究进展[ Advances on the strategies of soil phosphate solubilizing microorganisms to promote plant phosphorus uptake]. 生态学报 [Acta Ecologica Sinica] 43(11):4390−4399 (in Chinese)

doi: 10.5846/stxb202111193253

[46]

Chiaramonte JB. 2018. The rhizosphere microbiome of common bean (Phaseolus vulgaris L.) and the effects on phosphorus uptake. Doctoral thesis. Luiz de Queiroz College of Agriculture, Brazil. pp. 99−110 doi: 10.11606/T.11.2019.tde-17012019-161756

[47]

Jia LH, Wang X, Zhang R, Lin DL, Qiu R, et al. 2019. Rhizosphere microbial community mediated a synergism regulatory mechanism of phosphorus stress and immunity in plants by the integrated pathway. Journal of Plant Nutrition and Fertilizers 25(2):321−327

doi: 10.11674/zwyf.17441

[48]

Castrillo G, Teixeira PJPL, Paredes SH, Law TF, de Lorenzo L, et al. 2017. Root microbiota drive direct integration of phosphate stress and immunity. Nature 543:513–518

doi: 10.1038/s41586-026-10320-1

[49]

Jurburg SD, Salles JF. 2015. Functional redundancy and ecosystem function—the soil microbiota as a case study. In Biodiversity in Ecosystems - Linking Structure and Function, eds. Lo YH, Blanco JA, Roy S. pp. 29−42 doi: 10.5772/58981

[50]

Sauma-Sánchez T, Alcorta J, Tamayo-Leiva J, Díez B, Bezuidenhout H, et al. 2024. Functional redundancy buffers the effect of poly-extreme environmental conditions on southern African dryland soil microbial communities. FEMS Microbiology Ecology 100(12):fiae157

doi: 10.1093/femsec/fiae157

[51]

Jia Y, Whalen JK. 2020. A new perspective on functional redundancy and phylogenetic niche conservatism in soil microbial communities. Pedosphere 30(1):18−24

doi: 10.1016/S1002-0160(19)60826-X

[52]

Salazar A, Sulman BN, Dukes JS. 2018. Microbial dormancy promotes microbial biomass and respiration across pulses of drying-wetting stress. Soil Biology and Biochemistry 116:237−244

doi: 10.1016/j.soilbio.2017.10.017

[53]

Du H, Pan J, Zou D, Huang Y, Liu Y, et al. 2022. Microbial active functional modules derived from network analysis and metabolic interactions decipher the complex microbiome assembly in mangrove sediments. Microbiome 10(1):224

doi: 10.1186/s40168-022-01421-w

[54]

Trivedi P, Leach JE, Tringe SG, Sa T, Singh BK. 2020. Plant–microbiome interactions: from community assembly to plant health. Nature Reviews Microbiology 18(11):607−621

doi: 10.1038/s41579-020-0412-1

[55]

Radajewski S, Ineson P, Parekh NR, Murrell JC. 2000. Stable-isotope probing as a tool in microbial ecology. Nature 403(6770):646−649

doi: 10.1038/35001054

[56]

Alcolombri U, Pioli R, Stocker R, Berry D. 2022. Single-cell stable isotope probing in microbial ecology. ISME Communications 2(1):55

doi: 10.1038/s43705-022-00142-3

[57]

Aßhauer KP, Wemheuer B, Daniel R, Meinicke P. 2015. Tax4Fun: predicting functional profiles from metagenomic 16S rRNA data. Bioinformatics 31(17):2882−2884

doi: 10.1093/bioinformatics/btv287

[58]

Paul V, Banerjee Y, Ghosh P, Busi SB. 2020. Depthwise microbiome and isotopic profiling of a moderately saline microbial mat in a solar saltern. Scientific Reports 10(1):20686

doi: 10.1038/s41598-020-77622-w

[59]

Allison SD, Martiny JBH. 2008. Resistance, resilience, and redundancy in microbial communities. Proceedings of the National Academy of Sciences of the United States of America 105:11512−11519

doi: 10.1073/pnas.0801925105

[60]

Louca S, Polz MF, Mazel F, Albright MBN, Huber JA, et al. 2018. Function and functional redundancy in microbial systems. Nature Ecology & Evolution 2(6):936−943

doi: 10.1038/s41559-018-0519-1