[1]

Food and Agriculture Organization of the United Nations. 2022. Production of peaches and nectarines in 2022. Food and Agriculture Organization Statistics Division. https://www.fao.org/faostat/en/

[2]

Lipińska M, Tomaszewska M, Kołożyn-Krajewska D. 2019. Identifying factors associated with food losses during transportation: potentials for social purposes. Sustainability 11(7):2046

doi: 10.3390/su11072046
[3]

Makule E, Dimoso N, Tassou SA. 2022. Precooling and cold storage methods for fruits and vegetables in sub-Saharan Africa: a review. Horticulturae 8(9):776

doi: 10.3390/horticulturae8090776
[4]

Navarro A, Giménez R, Cantín CM, Martínez-García PJ, Val J, et al. 2022. Chilling injury in local and modern peach cultivars from a Spanish peach bank germplasm. Acta Horticulturae 1352(1):237−244

doi: 10.17660/ActaHortic.2022.1352.32
[5]

Lurie S. 2022. Proteomic and metabolomic studies on chilling injury in peach and nectarine. Frontiers in Plant Science 13:958312

doi: 10.3389/fpls.2022.958312
[6]

Goswami AK, Maurya NK, Goswami S, Bardhan K, Singh SK, et al. 2022. Physio-biochemical and molecular stress regulators and their crosstalk for low-temperature stress responses in fruit crops: a review. Frontiers in Plant Science 13:1022167

doi: 10.3389/fpls.2022.1022167
[7]

Muto A, Bruno L, Madeo ML, Ludlow R, Ferrari M, et al. 2022. Comparative transcriptomic profiling of peach and nectarine cultivars reveals cultivar-specific responses to chilled postharvest storage. Frontiers in Plant Science 13:1062194

doi: 10.3389/fpls.2022.1062194
[8]

Gohari G, Molaei S, Kheiry A, Ghafouri M, Razavi F, et al. 2021. Exogenous application of proline and L-cysteine alleviates internal browning and maintains eating quality of cold stored flat 'Maleki' peach fruits. Horticulturae 7(11):469

doi: 10.3390/horticulturae7110469
[9]

Bao Z, Zhou Q, Yu Y, Chen W, Yang Z, et al. 2024. Melatonin treatment induces DNA methylation to alleviate chilling induced-browning in cold stored peach fruit. Postharvest Biology and Technology 208:112686

doi: 10.1016/j.postharvbio.2023.112686
[10]

Liu Y, Wu J, Li Y, Deng W, Cao K, et al. 2024. Metabolism and transcriptional regulation in chilling injury development of nectarine fruit during postharvest cold storage. Postharvest Biology and Technology 210:112748

doi: 10.1016/j.postharvbio.2023.112748
[11]

Wang H, Zhang S, Wang Z, Li D, Yan L, et al. 2024. Resistance index and browning mechanism of apple peel under high temperature stress. Horticultural Plant Journal 10(2):305−317

doi: 10.1016/j.hpj.2022.10.013
[12]

Jiang Y, Fu J. 1998. Inhibition of polyphenol oxidase and the browning control of litchi fruit by glutathione and citric acid. Food Chemistry 62(1):49−52

doi: 10.1016/s0308-8146(97)00144-1
[13]

Wesam A, Northcutt JK, Dawson PD. 2023. Effect of pre-treatment and freezing on the polyphenol oxidase activity and color stability of sliced peaches. Journal of the Science of Food and Agriculture 103(7):3376−3389

doi: 10.1002/jsfa.12495
[14]

Liu B, Zhong R, Wei J, Zhang J, Luo H, et al. 2024. Genome-wide identification and analysis of the laccase gene family in Litchi chinensis Sonn. provides new insights into pericarp browning. Postharvest Biology and Technology 217:113108

doi: 10.1016/j.postharvbio.2024.113108
[15]

Liu B, Fang F, Guan H, Zhang J, Luo H, et al. 2024. Integrated function of proanthocyanidin and lignin polymerization mediated by LAC/PRXs in pericarp browning of Longan fruit. Postharvest Biology and Technology 207:112618

doi: 10.1016/j.postharvbio.2023.112618
[16]

Su J, Fu J, Wang Q, Silva C, Cavaco-Paulo A. 2018. Laccase: a green catalyst for the biosynthesis of poly-phenols. Critical Reviews in Biotechnology 38(3):294−307

doi: 10.1080/07388551.2017.1354353
[17]

Bai Y, Ali S, Liu S, Zhou J, Tang Y. 2023. Characterization of plant laccase genes and their functions. Gene 852:147060

doi: 10.1016/j.gene.2022.147060
[18]

Pourcel L, Routaboul JM, Kerhoas L, Caboche M, Lepiniec L, et al. 2005. TRANSPARENT TESTA10 encodes a laccase-like enzyme involved in oxidative polymerization of flavonoids in Arabidopsis seed coat. The Plant Cell 17(11):2966−2980

doi: 10.1105/tpc.105.035154
[19]

Gong Y, Song J, Du L, Vinqvist M, Palmer LC, Fillmore S, et al. 2018. Characterization of laccase from apple fruit during postharvest storage and its response to diphenylamine and 1-methylcyclopropene treatments. Food Chemistry 253:314−321

doi: 10.1016/j.foodchem.2018.01.142
[20]

Wang H, Zhang S, Fu Q, Wang Z, Liu X, et al. 2023. Transcriptomic and metabolomic analysis reveals a protein module involved in preharvest apple peel browning. Plant Physiology 192:2102−2122

doi: 10.1093/plphys/kiad064
[21]

Wei J, Zhang X, Zhong R, Liu B, Zhang X, et al. 2021. Laccase-mediated flavonoid polymerization leads to the pericarp browning of litchi fruit. Journal of Agricultural and Food Chemistry 69:15218−15230

doi: 10.1021/acs.jafc.1c06043
[22]

Tanou G, Minas IS, Scossa F, Belghazi M, Xanthopoulou A, et al. 2017. Exploring priming responses involved in peach fruit acclimation to cold stress. Scientific Reports 7:11358

doi: 10.1038/s41598-017-11933-3
[23]

Lurie S. 2021. Genomic and transcriptomic studies on chilling injury in peach and nectarine. Postharvest Biology and Technology 174:111444

doi: 10.1016/j.postharvbio.2020.111444
[24]

Dweh TJ, Kayastha S, Mahapatra M, Sahoo JP. 2023. Understanding the transcription factor mediated regulatory mechanism towards abiotic stress response in cereal crops. AgroEnvironmental Sustainability 1(3):265−273

doi: 10.59983/s2023010308
[25]

Viswanath KK, Kuo SY, Tu CW, Hsu YH, Huang YW, et al. 2023. The role of plant transcription factors in the fight against plant viruses. International Journal of Molecular Sciences 24:8433

doi: 10.3390/ijms24098433
[26]

Sun HJ, Luo ML, Zhou X, Zhou Q, Sun YY, et al. 2020. PuMYB21/PuMYB54 coordinate to activate PuPLDβ1 transcription during peel browning of cold-stored 'Nanguo' pears. Horticulture Research 7:136

doi: 10.1038/s41438-020-00356-3
[27]

Wang H, Cheng X, Yin D, Chen D, Luo C, et al. 2023. Advances in the research on plant WRKY transcription factors responsive to external stresses. Current Issues in Molecular Biology 45:2861−2880

doi: 10.3390/cimb45040187
[28]

Joshi R, Wani SH, Singh B, Bohra A, Dar ZA, et al. 2016. Transcription factors and plants response to drought stress: current understanding and future directions. Frontiers in Plant Science 7:1029

doi: 10.3389/fpls.2016.01029
[29]

Liu X, Song Y, Xing F, Wang N, Wen F, et al. 2016. GhWRKY25, a group I WRKY gene from cotton, confers differential tolerance to abiotic and biotic stresses in transgenic Nicotiana benthamiana. Protoplasma 253(5):1265−1281

doi: 10.1007/s00709-015-0885-3
[30]

Ma Z, Hu L. 2024. WRKY transcription factor responses and tolerance to abiotic stresses in plants. International Journal of Molecular Sciences 25:6845

doi: 10.3390/ijms25136845
[31]

Ye YJ, Xiao YY, Han YC, Shan W, Fan ZQ, et al. 2016. Banana fruit VQ motif-containing protein5 represses cold-responsive transcription factor MaWRKY26 involved in the regulation of JA biosynthetic genes. Scientific Reports 6:23632

doi: 10.1038/srep23632
[32]

Dan Y, Zhang S, Matherly A. 2016. Regulation of hydrogen peroxide accumulation and death of Agrobacterium-transformed cells in tomato transformation. Plant Cell, Tissue and Organ Culture (PCTOC) 127(1):229−236

doi: 10.1007/s11240-016-1045-y
[33]

Lin H, Bai L, Wei W, Su W, Wu Y, et al. 2024. The role of MaWRKY70 in regulating lipoxygenase gene transcription during chilling injury development in banana fruit. Foods 13(6):854

doi: 10.3390/foods13060854
[34]

Jin P, Zhu H, Wang L, Shan T, Zheng Y. 2014. Oxalic acid alleviates chilling injury in peach fruit by regulating energy metabolism and fatty acid contents. Food Chemistry 161:87−93

doi: 10.1016/j.foodchem.2014.03.103
[35]

Tatsuki M, Nakajima N, Fujii H, Shimada T, Nakano M, et al. 2013. Increased levels of IAA are required for system 2 ethylene synthesis causing fruit softening in peach (Prunus persica L. Batsch). Journal of Experimental Botany 64:1049−1059

doi: 10.1093/jxb/ers381
[36]

Livak KJ, Schmittgen TD. 2001. Analysis of relative gene expression data using real-time quantitative PCR and the 2−ΔΔCT method. Methods 25:402−408

doi: 10.1006/meth.2001.1262
[37]

Li SJ, Yin XR, Wang WL, Liu XF, Zhang B, et al. 2017. Citrus CitNAC62 cooperates with CitWRKY1 to participate in citric acid degradation via up-regulation of CitAco3. Journal of Experimental Botany 68:3419−3426

doi: 10.1093/jxb/erx187
[38]

Zhang AD, Wang WQ, Tong Y, Li MJ, Grierson D, et al. 2018. Transcriptome analysis identifies a zinc finger protein regulating starch degradation in kiwifruit. Plant Physiology 178:850−863

doi: 10.1104/pp.18.00427
[39]

Wang MM, Zhu QG, Deng CL, Luo ZR, Sun NJ, et al. 2017. Hypoxia-responsive ERFs involved in postdeastringency softening of persimmon fruit. Plant Biotechnology Journal 15:1409−1419

doi: 10.1111/pbi.12725
[40]

Wang Y, Deng L, Meng J, Niu L, Pan L, et al. 2021. Transcriptomic and metabolic analyses reveal the mechanism of ethylene production in stony hard peach fruit during cold storage. International Journal of Molecular Sciences 22:11308

doi: 10.3390/ijms222111308
[41]

Han YC, Fu CC. 2019. Cold-inducible MaC2H2s are associated with cold stress response of banana fruit via regulating MaICE1. Plant Cell Reports 38:673−680

doi: 10.1007/s00299-019-02399-w
[42]

Zhan W, Wang Y, Duan W, Li A, Miao Y, et al. 2024. Preliminary analysis, combined with omics, of chilling injury mechanism of peach fruits with different cold sensitivities during postharvest cold storage. Horticulturae 10(1):46

doi: 10.3390/horticulturae10010046
[43]

Khedr EH, Khedr N. 2023. Optimization of postharvest progesterone treatment to alleviate chilling injury in mango fruit, maintaining intracellular energy, cell wall stability, and antioxidant activity. Postharvest Biology and Technology 206:112572

doi: 10.1016/j.postharvbio.2023.112572
[44]

Wang J, Li F, Li B, Li L, Shang J, et al. 2025. ROS-mediated membrane damage and antioxidant imbalance drive apple flesh browning during cold storage. Frontiers in Plant Science 16:1718635

doi: 10.3389/fpls.2025.1718635
[45]

Lin Y, Jiang L, Chen Q, Li Y, Zhang Y, et al. 2018. Comparative transcriptome profiling analysis of red- and white-fleshed strawberry (Fragaria × ananassa) provides new insight into the regulation of the anthocyanin pathway. Plant & Cell Physiology 59:1844−1859

doi: 10.1093/pcp/pcy098
[46]

de la Rosa LA, Mercado-Mercado G, Rodrigo-García J, González-Aguilar GA, Alvarez-Parrilla E. 2010. Peach polyphenol oxidase inhibition by β-cyclodextrin and 4-hexylresorcinol is substrate dependent. CyTA − Journal of Food 8:87−93

doi: 10.1080/19476330903146013
[47]

Li F, Li Z, Gao Z, Wang G, Li H, et al. 2023. A laccase gene (LcLac) was involved in polyphenol metabolism and tissue browning of litchi callus. Scientia Horticulturae 321:112291

doi: 10.1016/j.scienta.2023.112291
[48]

Chen X, Zhang T, Wang H, Zhao W, Guo Z. 2025. Transcription factor WRKY complexes in plant signaling pathways. Phytopathology Research 7(1):54

doi: 10.1186/s42483-025-00349-x
[49]

Liu W, Liang X, Cai W, Wang H, Liu X, et al. 2022. Isolation and functional analysis of VvWRKY28, a Vitis vinifera WRKY transcription factor gene, with functions in tolerance to cold and salt stress in transgenic Arabidopsis thaliana. International Journal of Molecular Sciences 23:13418

doi: 10.3390/ijms232113418
[50]

Zhang Y, Yu H, Yang X, Li Q, Ling J, et al. 2016. CsWRKY46, a WRKY transcription factor from cucumber, confers cold resistance in transgenic-plant by regulating a set of cold-stress responsive genes in an ABA-dependent manner. Plant Physiology and Biochemistry 108:478−487

doi: 10.1016/j.plaphy.2016.08.013
[51]

Luo DL, Ba LJ, Shan W, Kuang JF, Lu WJ, et al. 2017. Involvement of WRKY transcription factors in abscisic-acid-induced cold tolerance of banana fruit. Journal of Agricultural and Food Chemistry 65:3627−3635

doi: 10.1021/acs.jafc.7b00915
[52]

Hou Y, Liu Y, Zhao L, Zhao Y, Wu Z, et al. 2023. EjCML19 and EjWRKY7 synergistically function in calcium chloride-alleviated chilling injury of loquat fruit. Postharvest Biology and Technology 203:112417

doi: 10.1016/j.postharvbio.2023.112417
[53]

Boro P, Sultana A, Mandal K, Chattopadhyay S. 2022. Interplay between glutathione and mitogen-activated protein kinase 3 via transcription factor WRKY40 under combined osmotic and cold stress in Arabidopsis. Journal of Plant Physiology 271:153664

doi: 10.1016/j.jplph.2022.153664
[54]

Xie Z, Zhang ZL, Zou X, Huang J, Ruas P, et al. 2005. Annotations and functional analyses of the rice WRKY gene superfamily reveal positive and negative regulators of abscisic acid signaling in aleurone cells. Plant Physiology 137:176−189

doi: 10.1104/pp.104.054312
[55]

Jiang L, Chen Y, Bi D, Cao Y, Tong J. 2021. Deciphering evolutionary dynamics of WRKY I genes in rosaceae species. Frontiers in Ecology and Evolution 9:801490

doi: 10.3389/fevo.2021.801490
[56]

Li S, Zhou X, Chen L, Huang W, Yu D. 2010. Functional characterization of Arabidopsis thaliana WRKY39 in heat stress. Molecules and Cells 29:475−484

doi: 10.1007/s10059-010-0059-2
[57]

Chen H, Lai Z, Shi J, Xiao Y, Chen Z, et al. 2010. Roles of Arabidopsis WRKY18, WRKY40, and WRKY60 transcription factors in plant responses to abscisic acid and abiotic stress. BMC Plant Biology 10:281

doi: 10.1186/1471-2229-10-281
[58]

Gao K, Zhou T, Hua Y, Guan C, Zhang Z. 2020. Transcription factor WRKY23 is involved in ammonium-induced repression of Arabidopsis primary root growth under ammonium toxicity. Plant Physiology and Biochemistry 150:90−98

doi: 10.1016/j.plaphy.2020.02.034
[59]

Yu Y, Wang L, Chen J, Liu Z, Park CM, et al. 2018. WRKY71 acts antagonistically against salt-delayed flowering in Arabidopsis thaliana. Plant & Cell Physiology 59(2):414−422

doi: 10.1093/pcp/pcx201