doi:10.48130/mycosphere-0026-0017

[1]

Wu F, Zhou LW, Yang ZL, Bau T, Li TH, et al. 2019. Resource diversity of Chinese macrofungi: edible, medicinal and poisonous species. Fungal Diversity 98(1):1−76

doi: 10.1007/s13225-019-00432-7

[2]

Hawksworth DL. 1991. The fungal dimension of biodiversity: magnitude, significance, and conservation. Mycological research 95(6):641−655

doi: 10.1016/S0953-7562(09)80810-1

[3]

Hibbett D, Nagy LG, Nilsson RH. 2025. Fungal diversity, evolution, and classification. Current Biology 35(11):R463−R469

doi: 10.1016/j.cub.2025.01.053

[4]

Hawksworth DL, Lücking R. 2017. Fungal diversity revisited: 2.2 to 3.8 million species. Microbiology Spectrum 5(4):0052

doi: 10.1128/microbiolspec.funk-0052-2016

[5]

Baldrian P. 2017. Forest microbiome: diversity, complexity and dynamics. FEMS Microbiology Reviews 41(2):109−130

doi: 10.1093/femsre/fuw040

[6]

Lundell TK, Mäkelä MR, Hildén K. 2010. Lignin-modifying enzymes in filamentous basidiomycetes—ecological, functional and phylogenetic review. Journal of Basic Microbiology 50(1):5−20

doi: 10.1002/jobm.200900338

[7]

Gadd GM. 2021. Fungal biomineralization. Current Biology 31(24):R1557−R1563

doi: 10.1016/j.cub.2021.10.041

[8]

Tedersoo L, Bahram M, Põlme S, Kõljalg U, Yorou NS, et al. 2014. Global diversity and geography of soil fungi. Science 346:1256688

doi: 10.1126/science.1256688

[9]

Hyde K, Tavakol Noorabadi M, Thiyagaraja V, He MQ, Johnston P, et al. 2024. The 2024 Outline of Fungi and fungus-like taxa. Mycosphere 15(1):5146−6239

doi: 10.5943/mycosphere/15/1/25

[10]

Zhao H, Yuan HS, Cui YJ, Wang K, Dai YC, et al. 2026. Global polypore diversity and distribution patterns. Fungal Diversity 136:136002

doi: 10.65390/fdiv.2026.136002

[11]

Tedersoo L, Bahram M, Zobel M. 2020. How mycorrhizal associations drive plant population and community biology. Science 367(6480):eaba1223

doi: 10.1126/science.aba1223

[12]

van der Heijden MGA, Martin FM, Selosse MA, Sanders IR. 2015. Mycorrhizal ecology and evolution: the past, the present, and the future. New Phytologist 205(4):1406−1423

doi: 10.1111/nph.13288

[13]

Steidinger BS, Crowther TW, Liang J, Van Nuland ME, Werner GDA, et al. 2019. Climatic controls of decomposition drive the global biogeography of forest-tree symbioses. Nature 569:404−408

doi: 10.1038/s41586-019-1128-0

[14]

Cai Q, Codjia JEI, Buyck B, Cui YY, Ryberg M, et al. 2024. The evolution of ectomycorrhizal symbiosis and host-plant switches are the main drivers for diversification of Amanitaceae (Agaricales, Basidiomycota). BMC Biology 22(1):230

doi: 10.1186/s12915-024-02031-8

[15]

Niego AGT, Rapior S, Thongklang N, Raspé O, Hyde KD, et al. 2023. Reviewing the contributions of macrofungi to forest ecosystem processes and services. Fungal Biology Reviews 44:100294

doi: 10.1016/j.fbr.2022.11.002

[16]

Ling ZL, Ma XB, Zhao RL, Cao B, Huang CY, et al. 2025. Overview of the Chinese edible fungi industry of and developing trends analysis. Mycosphere 16(1):2712−2775

doi: 10.5943/mycosphere/16/1/17

[17]

Manawasinghe I, Hyde K, Balasuriya A, Suwannarach N, Boonyuen N, et al. 2025. The emerging role of Fungi in sustainable farming and global food security. Mycosphere 16(1):4936−5064

doi: 10.5943/mycosphere/16/1/37

[18]

Niego AGT, Lambert C, Mortimer P, Thongklang N, Rapior S, et al. 2023. The contribution of fungi to the global economy. Fungal Diversity 121(1):95−137

doi: 10.1007/s13225-023-00520-9

[19]

Song CG, Xu TM, Xu YH, Wang D, Long Z, et al. 2025. Systematic revision, molecular phylogeny and divergence times of Thelephorales (Basidiomycota). Mycosphere 16(1):296−422

doi: 10.5943/mycosphere/16/1/5

[20]

Habib K, Li W, Ren Y, Liu L, Lu C, et al. 2025. Exploration of ascomycetous fungi revealing novel taxa in Southwestern China. Mycosphere 16(1):1412−1529

doi: 10.5943/mycosphere/16/1/9

[21]

Dong JH, Chen ML, Chen M, Li Q, Zhu YJ, et al. 2025. Notes, outline, taxonomy and phylogeny of wood-inhabiting Agaricales. Mycosphere 16(1):2599−2711

doi: 10.5943/mycosphere/16/1/16

[22]

Antonelli A, Fry C, Smith RJ, Eden J, Kersey PJ, et al. 2020. State of the World's plants and fungi 2026. doi: 10.34885/172

[23]

Varga T, Krizsán K, Földi C, Dima B, Sánchez-García M, et al. 2019. Megaphylogeny resolves global patterns of mushroom evolution. Nature Ecology & Evolution 3:668−678

doi: 10.1038/s41559-019-0834-1

[24]

Nagy LG, Branco S, Floudas D, Hibbett DS, Lofgren L, et al. 2026. The biodiversity, genomics, ecology and evolution of mushroom-forming fungi. Nature Reviews Biodiversity 2:24−39

doi: 10.1038/s44358-025-00107-z

[25]

Tedersoo L, Mikryukov V, Zizka A, Bahram M, Hagh-Doust N, et al. 2022. Global patterns in endemicity and vulnerability of soil fungi. Glob Chang Biol 28:6696−6710

doi: 10.1111/gcb.16398

[26]

Barbi F, Martinović T, Odriozola I, Machac A, Moravcová A, et al. 2025. Disentangling drivers behind fungal diversity gradients along altitude and latitude. New Phytologist 247(1):295−308

doi: 10.1111/nph.70012

[27]

Wang XH, Cai Q, Yu FM, Yang ZL, Zhou SY, et al. 2025. New and notable taxa of Basidiomycota on the Qinghai-Xizang Plateau and its surrounding areas. Fungal Diversity 133(1):235−437

doi: 10.1007/s13225-025-00558-x

[28]

Singer R. 1986. The Agaricales in modern taxonomy. 4th ed. Koenigstein, Germany: Koeltz Scientific Books. pp. 1–981

[29]

Largent DL. 1986. How to identify mushrooms to genus I: macroscopic features. Eureka, California: Mad River Press. pp. 1–166

[30]

Largent DL, Johnson D, Watling R. 1977. How to identify mushrooms to genus III: microscopic features. Eureka, California: Mad River Press. pp. 1–148

[31]

Liu F, Wang SH, Cheewangkoon R, Zhao RL. 2024. Uneven distribution of prokaryote-derived horizontal gene transfer in fungi: a lifestyle-dependent phenomenon. mBio 16(1):e02855-24

doi: 10.1128/mbio.02855-24

[32]

Alster CJ, Allison SD, Johnson NG, Glassman SI, Treseder KK. 2021. Phenotypic plasticity of fungal traits in response to moisture and temperature. ISME Communications 1:43

doi: 10.1038/s43705-021-00045-9

[33]

Wrzosek M, Ruszkiewicz-Michalska M, Sikora K, Damszel M, Sierota Z. 2017. The plasticity of fungal interactions. Mycological Progress 16(2):101−108

doi: 10.1007/s11557-016-1257-x

[34]

Lin P, Kook M, Yi TH, Yan ZF. 2023. Current fungal taxonomy and developments in the identification system. Current Microbiology 80(12):375

doi: 10.1007/s00284-023-03514-7

[35]

Sánchez-García M, Ryberg M, Khan FK, Varga T, Nagy LG, et al. 2020. Fruiting body form, not nutritional mode, is the major driver of diversification in mushroom-forming fungi. Proceedings of the National Academy of Sciences of the United States of America 117(51):32528−32534

doi: 10.1073/pnas.1922539117

[36]

Virágh M, Merényi Z, Csernetics Á, Földi C, Sahu N, et al. 2021. Evolutionary morphogenesis of sexual fruiting bodies in Basidiomycota: toward a new evo-devo synthesis. Microbiology and Molecular Biology Reviews 86(1):e00019-21

doi: 10.1128/MMBR.00019-21

[37]

Lücking R, Aime MC, Robbertse B, Miller AN, Aoki T, et al. 2021. Fungal taxonomy and sequence-based nomenclature. Nature Microbiology 6:540−548

doi: 10.1038/s41564-021-00888-x

[38]

de Queiroz K, Gauthier J. 1990. Phylogeny as a central principle in taxonomy: phylogenetic definitions of taxon names. Systematic Biology 39(4):307−322

doi: 10.2307/2992353

[39]

Tekpinar AD, Kalmer A. 2019. Utility of various molecular markers in fungal identification and phylogeny. Nova Hedwigia 109(1–2):187−224

doi: 10.1127/nova_hedwigia/2019/0528

[40]

Wiens JJ, Blomenkamp EG, Corliss M, Li X. 2026. Global biodiversity and the expanding tree of life. Proceedings of the National Academy of Sciences of the United States of America 123(12):e2530656123

doi: 10.1073/pnas.2530656123

[41]

Ke W, Lei C. 2023. 中国菌物分类学和多样性研究的历史与现状概况[An overview of the historical and current status of fungal taxonomy and diversity in China]. 菌物学报[Acta Mycologica Sinica] 42(1):50−62 (in Chinese)

doi: 10.13346/j.mycosystema.220497

[42]

Mittermeier R, Gil PR, Hoffmann M, Pilgrim J, Brooks T, et al. 2004. Hotspots revisited: Earth's biologically richest and most endangered terrestrial ecoregions. Mexico City, Mexico: CEMEX

[43]

Yao T, Thompson LG, Mosbrugger V, Zhang F, Ma Y, et al. 2012. Third pole environment (TPE). Environmental Development 3:52−64

doi: 10.1016/j.envdev.2012.04.002

[44]

Mosbrugger V, Favre A, Muellner-Riehl A, Päckert M, Mulch A. 2018. Cenozoic evolution of geo-biodiversity in the Tibeto-Himalayan region. In Mountains, Climate, and Biodiversity, eds. Hoorn C, Perrigo A, Antonelli A. 1st ed. Hoboken, NJ: Wiley-Blackwell. pp. 429–449

[45]

Yang ZL. 2005. Diversity and biogeography of higher fungi in China. In Evolutionary genetics of fungi. Norfolk, UK: Horizon Bioscience. pp. 35–62

[46]

Tapponnier P, Zhiqin X, Roger F, Meyer B, Arnaud N, et al. 2001. Oblique stepwise rise and growth of the Tibet plateau. Science 294(5547):1671−1677

doi: 10.1126/science.105978

[47]

Feng B, Yang Z. 2018. Studies on diversity of higher fungi in Yunnan, southwestern China: a review. Plant Diversity 40(4):165−171

doi: 10.1016/j.pld.2018.07.001

[48]

Han XX, Phurbu D, Cao B, Li JX, Zhu XY, et al. 2025. Molecular phylogeny and morphology reveal four new species of Conocybe (Bolbitiaceae, Agaricales) from the Qinghai–Xizang Plateau, China. Journal of Fungi 11(1):45

doi: 10.3390/jof11010045

[49]

Naranjo-Ortiz MA, Gabaldón T. 2019. Fungal evolution: major ecological adaptations and evolutionary transitions. Biological Reviews 94(4):1443−1476

doi: 10.1111/brv.12510

[50]

Körner C. 1995. Alpine plant diversity: a global survey and functional interpretations. In Arctic and Alpine Biodiversity: Patterns, Causes and Ecosystem Consequences, eds. Chapin FS, Körner C. Ecological Studies. Vol. 113. Berlin, Heidelberg: Springer. pp. 45–62 doi: 10.1007/978-3-642-78966-3_4

[51]

Shooner S, Davies TJ, Saikia P, Deka J, Bharali S, et al. 2018. Phylogenetic diversity patterns in Himalayan forests reveal evidence for environmental filtering of distinct lineages. Ecosphere 9(5):e02157

doi: 10.1002/ecs2.2157

[52]

Peay KG, Kennedy PG, Talbot JM. 2016. Dimensions of biodiversity in the Earth mycobiome. Nature Reviews Microbiology 14:434−447

doi: 10.1038/nrmicro.2016.59

[53]

Shen C, Gunina A, Luo Y, Wang J, He JZ, et al. 2020. Contrasting patterns and drivers of soil bacterial and fungal diversity across a mountain gradient. Environmental Microbiology 22(8):3287−3301

doi: 10.1111/1462-2920.15090

[54]

Vetaas OR. 2021. Mountain biodiversity and elevational gradients. Frontiers of Biogeography 13(3):e54146

doi: 10.21425/F5FBG54146

[55]

Tenorio EA, Montoya P, Norden N, Rodríguez-Buriticá S, Salgado-Negret B, et al. 2023. Mountains exhibit a stronger latitudinal diversity gradient than lowland regions. Journal of Biogeography 50(6):1026−1036

doi: 10.1111/jbi.14597

[56]

Costa FVd, Viana-Júnior AB, Aguilar R, Silveira FAO, Cornelissen TG. 2023. Biodiversity and elevation gradients: insights on sampling biases across worldwide mountains. Journal of Biogeography 50(11):1879−1889

doi: 10.1111/jbi.14696

[57]

Antonelli A, Kissling WD, Flantua SGA, Bermúdez MA, Mulch A, et al. 2018. Geological and climatic influences on mountain biodiversity. Nature Geoscience 11:718−725

doi: 10.1038/s41561-018-0236-z

[58]

Perrigo A, Hoorn C, Antonelli A. 2020. Why mountains matter for biodiversity. Journal of Biogeography 47(2):315−325

doi: 10.1111/jbi.13731

[59]

Cui YJ, Wu YD, Jiang YH, Zhu AH, Wu F, et al. 2025. Diversity of macrofungi in southeast Xizang 1. The wood-decay fungi. Mycology 16(2):635−669

doi: 10.1080/21501203.2024.2379476

[60]

Liu X, Chen Q, Xiang Q, Gu Y, Han X, et al. 2025. Additions to the Entoloma (Agaricales, Entolomataceae) from China: description of five species with one new to science. MycoKeys 123:89−103

doi: 10.3897/mycokeys.123.162577

[61]

Wang K, Liu SL, Liu XZ, Hong P, Wei HW, et al. 2024. Catalogue of fungi in China 3. New taxa of macrofungi from southern Xizang, China. Mycology 16:91−123

doi: 10.1080/21501203.2024.2392014

[62]

Xu R, Hyde K, Li JN, Boonmee S, Liu NG, et al. 2025. Lignicolous freshwater fungi of the pan Qinghai-Xizang Plateau, China. Fungal Diversity 133(1):23−234

doi: 10.1007/s13225-025-00555-0

[63]

Zhu Y, Zhou L, Liu J, Yuan Y, Wei Y, et al. 2026. Thirty new species in Thelephorales and Hymenochaetales from the Eastern Himalayas and Southern China. Mycosphere 17:e001

doi: 10.48130/mycosphere-0026-0001

[64]

Liu S, Pan M, Cui BK, Zhu B. 2025. Catalogue of fungi in China 5. Preliminary survey of macrofungi in Medog, Southwest China. Mycology 16(4):1589−1636

doi: 10.1080/21501203.2025.2471382

[65]

Wang SH, Li GJ, Phurbu D, He MQ, Zhang MZ, et al. 2024. Four new species of Russula from the Xizang Autonomous Region and other provinces of China. Mycology 15(2):210−237

doi: 10.1080/21501203.2023.2265667

[66]

Li JX, Cao B, Phurbu D, He MQ, Zhu XY, et al. 2024. The revision of the taxonomic system of Lycoperdaceae. Mycosphere 15(1):4919−5016

doi: 10.5943/mycosphere/15/1/22

[67]

Xie ML, Dima B, Wang K, Phukhamsakda C, Li Y, et al. 2025. Taxonomy and phylogeny of Cortinarius sect. Anomali in China. Persoonia 54:225−263

doi: 10.3114/persoonia.2025.54.07

[68]

Zhou Y, Jia X, Han L, Liu Z, Kang S, et al. 2021. Fungal community diversity in soils along an elevation gradient in a Quercus aliena var. acuteserrata forest in Qinling Mountains, China. Applied Soil Ecology 167:104104

doi: 10.1016/j.apsoil.2021.104104

[69]

Zhang Y, Zhou DQ, Zhao Q, Zhou TX, Hyde KD. 2010. Diversity and ecological distribution of macrofungi in the Laojun Mountain region, southwestern China. Biodiversity and Conservation 19(12):3545−3563

doi: 10.1007/s10531-010-9915-9

[70]

Kornerup A, Wanscher J. 1967. Methuen handbook of colour. London, UK: Eyre Methuen

[71]

Tang SM, Vadthanarat S, Raghoonundon B, Luo ZL, Zhu XY, et al. 2024. New species and new records of Laccaria (Agaricales, Basidiomycota) from Northern Thailand. MycoKeys 107:189–217

doi: 10.3897/mycokeys.107.127907

[72]

Antonín V, Ďuriška O, Jančovičová S, Para R, Kudláček T, et al. 2022. Multilocus phylogeny and taxonomy of European Melanoleuca subgenus Melanoleuca. Mycologia 114(1):114−143

doi: 10.1080/00275514.2021.1966246

[73]

White TJ, Bruns T, Lee S, Taylor J. 1990. Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. In PCR protocols: a guide to methods and applications, eds. Innis MA, Gelfand DH, Sninsky JJ, White TJ. Cambridge, UK: Academic Press. pp. 315−322 doi: 10.1016/b978-0-12-372180-8.50042-1

[74]

Gardes M, Bruns TD. 1993. ITS primers with enhanced specificity for basidiomycetes—application to the identification of mycorrhizae and rusts. Molecular Ecology 2(2):113−118

doi: 10.1111/j.1365-294X.1993.tb00005.x

[75]

Hopple Jr JS, Vilgalys R. 1999. Phylogenetic relationships in the mushroom genus Coprinus and dark-spored allies based on sequence data from the nuclear gene coding for the large ribosomal subunit RNA: divergent domains, outgroups, and monophyly. Molecular Phylogenetics and Evolution 13(1):1−19

doi: 10.1006/mpev.1999.0634

[76]

Moncalvo JM, Lutzoni FM, Rehner SA, Johnson J, Vilgalys R. 2000. Phylogenetic relationships of agaric fungi based on nuclear large subunit ribosomal DNA sequences. Systematic Biology 49(2):278−305

doi: 10.1093/sysbio/49.2.278

[77]

Matheny PB. 2005. Improving phylogenetic inference of mushrooms with RPB1 and RPB2 nucleotide sequences (Inocybe; Agaricales). Molecular Phylogenetics and Evolution 35(1):1−20

doi: 10.1016/j.ympev.2004.11.014

[78]

Rehner SA, Buckley E. 2005. A Beauveria phylogeny inferred from nuclear ITS and EF1-α sequences: evidence for cryptic diversification and links to Cordyceps teleomorphs. Mycologia 97(1):84−98

doi: 10.1080/15572536.2006.11832842

[79]

Antonín V, Ďuriška O, Jančovičová S, Tomšovský M. 2015. Identity of Agaricus brevipes Bull. (Melanoleuca brevipes, Tricholomataceae, Basidiomycota). Mycological Progress 14(11):107

doi: 10.1007/s11557-015-1125-0

[80]

Xu YD, Zhang P, Chen ZH, He ZM. 2025. Three new species and two new records of the genus Laccaria (Agaricales, Basidiomycota) from subtropical China based on morphological and multi-locus phylogenetic evidence. MycoKeys 123:147−170

doi: 10.3897/mycokeys.123.156526

[81]

Long X, He B, Fang Y, Tang C. 2016. Identification and characterization of the glucose-6-phosphate dehydrogenase gene family in the para rubber tree, Hevea brasiliensis. Frontiers in Plant Science 7:215

doi: 10.3389/fpls.2016.00215

[82]

Katoh K, Rozewicki J, Yamada KD. 2017. MAFFT online service: multiple sequence alignment, interactive sequence choice and visualization. Briefings in Bioinformatics 20(4):1160−1166

doi: 10.1093/bib/bbx108

[83]

Hall TA. 1999. BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic Acids Symposium Series. Vol. 41. pp. 95–98 doi: 10.14601/Phytopathol_Mediterr-14998u1.29

[84]

Zhang D, Gao F, Jakovlić I, Zou H, Zhang J, et al. 2020. PhyloSuite: an integrated and scalable desktop platform for streamlined molecular sequence data management and evolutionary phylogenetics studies. Molecular Ecology Resources 20(1):348−355

doi: 10.1111/1755-0998.13096

[85]

Miller MA, Pfeiffer W, Schwartz T. 2010. Creating the CIPRES Science Gateway for inference of large phylogenetic trees. 2010 Gateway Computing Environments Workshop (GCE), New Orleans, LA, USA, 14 November 2010. New York, NY: IEEE doi: 10.1109/GCE.2010.5676129

[86]

Stamatakis A. 2006. RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models. Bioinformatics 22(21):2688−2690

doi: 10.1093/bioinformatics/btl446

[87]

Lanfear R, Frandsen PB, Wright AM, Senfeld T, Calcott B. 2017. PartitionFinder 2: new methods for selecting partitioned models of evolution for molecular and morphological phylogenetic analyses. Molecular Biology and Evolution 34(3):772−773

doi: 10.1093/molbev/msw260

[88]

Ronquist F, Teslenko M, van der Mark P, Ayres DL, Darling A, et al. 2012. MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology 61(3):539−542

doi: 10.1093/sysbio/sys029

[89]

Xie J, Chen Y, Cai G, Cai R, Hu Z, et al. 2023. Tree Visualization By One Table (tvBOT): a web application for visualizing, modifying and annotating phylogenetic trees. Nucleic Acids Research 51(W1):W587−W592

doi: 10.1093/nar/gkad359

[90]

Zhao RL, Zhou JL, Chen J, Margaritescu S, Sánchez-Ramírez S, et al. 2016. Towards standardizing taxonomic ranks using divergence times—a case study for reconstruction of the Agaricus taxonomic system. Fungal Diversity 78(1):239−292

doi: 10.1007/s13225-016-0357-x

[91]

Liang YS, Huang XX, Lin ZJ, He DL, Qiu LH. 2024. Three new species of Agaricus from Baiyun Mountain, Guangzhou, China. Mycologia 116(3):431−448

doi: 10.1080/00275514.2024.2311039

[92]

He MQ, Zhao RL, Liu DM, Denchev TT, Begerow D, et al. 2022. Species diversity of Basidiomycota. Fungal Diversity 114(1):281−325

doi: 10.1007/s13225-021-00497-3

[93]

Lin C, Liu B, Zhao M, Ma K, Ji L. 2025. Catalogue of life China: towards an index of known species present in China. The Innovation Life 3(3):100140

doi: 10.59717/j.xinn-life.2025.100141

[94]

Zhao R, Karunarathna S, Raspé O, Parra LA, Guinberteau J, et al. 2011. Major clades in tropical Agaricus. Fungal Diversity 51(1):279−296

doi: 10.1007/s13225-011-0136-7

[95]

Chen J, Callac P, Parra LA, Karunarathna SC, He MQ, et al. 2017. Study in Agaricus subgenus Minores and allied clades reveals a new American subgenus and contrasting phylogenetic patterns in Europe and Greater Mekong Subregion. Persoonia – Molecular Phylogeny and Evolution of Fungi 38:170−196

doi: 10.3767/003158517X695521

[96]

Parra LA, Angelini C, Miller KO, Chen J. 2024. Agaricus subiculosus, a new species of the genus Agaricus sect. Minores from Puerto Rico (USA). MycolObs - Mycological Observations 8:33−43

[97]

Cao B, Phurbu D, Ralaiveloarisoa A, Liimatainen K, Niskanen T, et al. 2025. Fungal diversity notes 1919–2016: taxonomic and phylogenetic contributions to fungal taxa. Fungal Diversity 132:425−651

doi: 10.1007/s13225-025-00556-z

[98]

Hussain S, Al-Kharousi M, Al-Muharabi MA, Al-Maqbali Da, Al-Shabibi Z, et al. 2022. Phylogeny of Agaricus subgenus Pseudochitonia with the description of a new section and a new species from Oman. Mycological Progress 21(8):72

doi: 10.1007/s11557-022-01819-8

[99]

Parra LA, Angelini C, Ortiz-Santana B, Mata G, Billette C, et al. 2018. The genus Agaricus in the Caribbean. Nine new taxa mostly based on collections from the Dominican Republic. Phytotaxa 345(3):219−271

doi: 10.11646/phytotaxa.345.3.2

[100]

He MQ, Chuankid B, Hyde KD, Cheewangkoon R, Zhao RL. 2018. A new section and species of Agaricus subgenus Pseudochitonia from Thailand. MycoKeys 40:53−67

doi: 10.3897/mycokeys.40.26918

[101]

Liu AQ, Dai RC, Zhang MZ, Cao B, Xi YL, et al. 2020. Species of Agaricus section Agaricus from China. Phytotaxa 452(1):1−18

doi: 10.11646/phytotaxa.452.1.1

[102]

Parra Sánchez LA. 2008. Fungi Europaei Agaricus L. Allopsalliota, Nauta & Bas. Fungi Europaei. Vol. 1. Alassio, Italy: Edizioni Candusso

[103]

Kerrigan RW. 2016. Agaricus of North America. New York: New York Botanical Garden Press

[104]

Liu AQ. 2020. 中国主要林区蘑菇属系统学研究[Phylogenetic study of Agaricus in main forests of China]. Thesis. 中国科学院大学[University of Chinese Academy of Sciences], Bejing, China (in Chinese). pp. 1–269

[105]

Yang KL, Lin JY, Li GM, Yang ZL. 2025. New and interesting fungi from Guangzhou City, South China. Phytotaxa 707(1):1−95

doi: 10.11646/phytotaxa.707.1.1

[106]

Wang SE, Bau T. 2024. Six new species of Agaricus (Agaricaceae, Agaricales) from Northeast China. Journal of Fungi 10(1):59

doi: 10.3390/jof10010059

[107]

Wang SE, Chen SA, Huang HC, Lin DM, Liu PH. 2025. Three new species of Agaricus (Agaricaceae, Agaricales) from southern China. MycoKeys 119:47−66

doi: 10.3897/mycokeys.119.154278

[108]

Cao B, He MQ, Ling ZL, Zhang MZ, Wei SL, et al. 2021. A revision of Agaricus section Arvenses with nine new species from China. Mycologia 113(1):191−211

doi: 10.1080/00275514.2020.1830247

[109]

He ZM, Chen ZH, Bau T, Wang GS, Yang ZL. 2023. Systematic arrangement within the family Clitocybaceae (Tricholomatineae, Agaricales): phylogenetic and phylogenomic evidence, morphological data and muscarine-producing innovation. Fungal Diversity 123(1):1−47

doi: 10.1007/s13225-023-00527-2

[110]

Qi Y, Xu A, Yang L, Guo H, Guo Y, et al. 2025. Multiple analyses reveal evidence for three new species of Collybia (Clitocybaceae, Basidiomycete) from China. Journal of Fungi 11(5):371

doi: 10.3390/jof11050371

[111]

Breitenbach J, Kränzlin F. 1991. Fungi of Switzerland 3: Boletales and Agaricales (Bolets and Agarics Pt. 1). Luzern: Mykologia

[112]

Ronikier M, Miśkiewicz A. 2002. Clitocybe dryadicola (J. Favre) Harmaja—a new species for Poland and the Carpathians recorded in Tatra mountains. Cryptogamie Mycologie 23(2):163−166

[113]

He MQ, Zhao RL, Hyde KD, Begerow D, Kemler M, et al. 2019. Notes, outline and divergence times of Basidiomycota. Fungal Diversity 99(1):105−367

doi: 10.1007/s13225-019-00435-4

[114]

Tang SM, Zhao G, Niu KY, Li RY, Yu FM, et al. 2025. Species Diversity of Edible Mushrooms I—four new Laccaria species from Yunnan Province, China. Journal of Fungi 11(3):189

doi: 10.3390/jof11030189

[115]

Li J, Che NJ, Cui YY. 2024. Three new species of Laccaria (Agaricales, Basidiomycota) from Southwest China (Yunnan) based on morphological and multi-gene sequence data. Frontiers in Microbiology 15:1411488

doi: 10.3389/fmicb.2024.1411488

[116]

Zhang SF, Gui Y, Zhu GS, Shang NJ, Li B, et al. 2024. Laccaria guizhouensis sp. nov. (Agaricales, Basidiomycota) from southwest China. New Zealand Journal of Botany 62(2–3):138−150

doi: 10.1080/0028825X.2024.2304754

[117]

Singer R. 1962. The Agaricales in modern taxonomy. 2nd ed. Weinheim: J Cramer

[118]

Berkeley MJ, Broome CE. 1883. XLIII.—notices of British fungi. Annals and Magazine of Natural History 12(72):370−374

doi: 10.1080/00222938309459647

[119]

Mueller GM. 1984. New North American species of Laccaria (Agaricales). Mycotaxon 20(1):101−116

doi: 10.5962/p.418792

[120]

Mueller GM, Vellinga EC. 1986. Taxonomic and nomenclatural notes on Laccaria B. & Br. Laccaria amethystea, L. fraterna, L. laccata, L. pumila, and their synonyms. Persoonia – Molecular Phylogeny and Evolution of Fungi 13:27−43

[121]

Mueller GM. 1991. Laccaria laccata complex in North America and Sweden: intercollection pairing and morphometric analyses. Mycologia 83(5):578−594

doi: 10.2307/3760213

[122]

Wilson AW, May TW, Mueller GM. 2017. Biogeography of the ectomycorrhizal mushroom genus Laccaria. In Biogeography of Mycorrhizal Symbiosis, ed. L Tedersoo. Ecological Studies. Cham: Springer International Publishing. pp. 273–297 doi: 10.1007/978-3-319-56363-3_13

[123]

Quan L, Shi L, Zhang S, Yao Q, Yang Q, et al. 2023. Ectomycorrhizal fungi, two species of Laccaria, differentially block the migration and accumulation of cadmium and copper in Pinus densiflora. Chemosphere 334:138857

doi: 10.1016/j.chemosphere.2023.138857

[124]

Cullings K, Desimone J. 2020. Rapid response by the Ectomycorrhizal (ECM) community of a lodgepole pine forest to diesel application as indicated by Laccase gene fragment diversity. Journal of Bioremediation & Biodegradation 5:235

doi: 10.4172/2155-6199.1000235

[125]

Hongsanan S, Khuna S, Manawasinghe IS, Tibpromma S, Chethana KWT, et al. 2025. Mycosphere notes 521–571: a special edition of fungal biodiversity to celebrate Kevin D. Hyde's 70th birthday and his exceptional contributions to Mycology. Mycosphere 16(2):2002–2180

doi: 10.5943/mycosphere/16/2/1

[126]

Zhang M, Gao XL, Mu LQ, Deng WQ. 2023. Morphology and molecular phylogeny reveal five new species of Laccaria (Hydnangiaceae, Agaricales) from Southern China. Journal of Fungi 9(12):1179

doi: 10.3390/jof9121179

[127]

Popa F, Rexer KH, Donges K, Yang ZL, Kost GW. 2014. Three new Laccaria species from Southwest China (Yunnan). Mycological Progress 13(4):1105–1117

doi: 10.1007/s11557-014-0998-7

[128]

Wilson AW, Hosaka K, Perry BA, Mueller GM. 2013. Laccaria (Agaricomycetes, Basidiomycota) from Tibet (Xizang Autonomous Region, China). Mycoscience 54(6):406−419

doi: 10.1016/j.myc.2013.01.006

[129]

Cui YY, Cai Q, Li J, Yang ZL. 2021. Two new Laccaria species from China based on molecular and morphological evidence. Mycological Progress 20(4):567−576

doi: 10.1007/s11557-021-01698-5

[130]

Cho HJ, Park MS, Lee H, Oh SY, Wilson AW, et al. 2018. A systematic revision of the ectomycorrhizal genus Laccaria from Korea. Mycologia 110(5):948−961

doi: 10.1080/00275514.2018.1507542

[131]

Dovana F, Scali E, Lopez Del Visco C, Moreno G, Para R, et al. 2025. Solving the enigma of the identity of Laccaria laccata. Journal of Fungi 11(8):575

doi: 10.3390/jof11080575

[132]

Orton PD. 1960. New check list of British Agarics and Boleti: part III. Notes on genera and species in the list. Transactions of the British Mycological Society 43(2):159−384

doi: 10.1016/S0007-1536(60)80065-4

[133]

Imai S. 1938. Studies on the Agaricaceae of Hokkaido I. Journal of the Faculty of Agriculture, Hokkaido Imperial University 43(1):1−178

[134]

Choudhary S, Pal Sharma Y, Uniyal P. 2025. Laccaria friesii sp. nov. (Hydnangiaceae), a new species from northwestern Himalaya of Uttarakhand, India. Nordic Journal of Botany 2025(5):e04617

doi: 10.1111/njb.04617

[135]

Li F. 2020. Two new species of Laccaria from South China, with a note on Hodophilus glaberipes. Mycological Progress 19(5):525−539

doi: 10.1007/s11557-020-01573-9

[136]

Mueller G. 1991. Laccaria longipes, a new North American species of the Laccaria laccata complex. Mycotaxon 40:145−150

doi: 10.5962/p.416080

[137]

McNabb RFR. 1972. The Tricholomataceae of New Zealand 1. Laccaria Berk. & Br. New Zealand Journal of Botany 10(3):461−484

doi: 10.1080/0028825X.1972.10428618

[138]

Ye L, Mortimer PE, Tang SM, Tao Q, He G, et al. 2025. Cuphophyllus lijiangensis sp. nov. (Hygrophoraceae, Agaricales) from Yunnan, China. New Zealand Journal of Botany 63(2–3):354−369

doi: 10.1080/0028825X.2024.2445273

[139]

Latha KPD, Varsha MK, Abhimanyu C, Ajmal M. 2025. Cuphophyllus aureus—a new species from Kerala state, India. Phytotaxa 721:265−276

doi: 10.11646/phytotaxa.721.3.6

[140]

Crous PW, Catcheside DEA, Catcheside PS, Alfenas AC, Alfenas RF, et al. 2025. Fungal Planet description sheets: 1781−1866. Persoonia – Molecular Phylogeny and Evolution of Fungi 54:327−587

doi: 10.3114/persoonia.2025.54.10

[141]

Xia W, Wang C, Li T, Deng W, Yue H, Zhang M. 2025. Cuphophyllus striatus (Cuphophyllaceae, Agaricales), a new species from southwestern China. Phytotaxa 701(2):207−215

doi: 10.11646/phytotaxa.701.2.7

[142]

Lodge DJ, Padamsee M, Matheny PB, Aime MC, Cantrell SA, et al. 2014. Molecular phylogeny, morphology, pigment chemistry and ecology in Hygrophoraceae (Agaricales). Fungal Diversity 64(1):1−99

doi: 10.1007/s13225-013-0259-0

[143]

Voitk A, Saar I, Lodge DJ, Boertmann D, Berch SM, et al. 2020. New species and reports of Cuphophyllus from northern North America compared with related Eurasian species. Mycologia 112(2):438−452

doi: 10.1080/00275514.2019.1703476

[144]

Griffith GW, Easton GL, Jones AW. 2002. Ecology and diversity of waxcap (Hygrocybe spp.) Fungi. Botanical Journal of Scotland 54(1):7−22

doi: 10.1080/03746600208685025

[145]

Bon M. 1984. Le genre Cuphophyllus (Donk) sp. nov. Documents Mycologiques 14(56):9−12

[146]

Boertmann D. 2010. The genus Hygrocybe. 2nd revised edition. Fungi of Northern Europe. Vol. 1. Denmark: Svampetryk. pp. 200

[147]

Zhang YX, Mao N, Li T, Fan L. 2024. Two new species of Infundibulicybe (Omphalinaceae, Agaricales) from northern China. Phytotaxa 649(2):193−210

doi: 10.11646/phytotaxa.649.2.4

[148]

Harmaja H. 2003. Notes on Clitocybe s. lato (Agaricales). Annales Botanici Fennici 40(3):213−218

[149]

Vizzini A, Contu M, Musumeci E, Ercole E. 2011. A new taxon in the Infundibulicybe gibba complex (Basidiomycota, Agaricales, Tricholomataceae) from Sardinia (Italy). Mycologia 103(1):203−208

doi: 10.3852/10-137

[150]

Vizzini A, Alvarado P, Consiglio G, Marchetti M, Xu J. 2024. Family matters inside the order Agaricales: systematic reorganization and classification of incertae sedis clitocyboid, pleurotoid and tricholomatoid taxa based on an updated 6-gene phylogeny. Studies in Mycology 107:67−148

doi: 10.3114/sim.2024.107.02

[151]

Binder M, Larsson KH, Matheny PB, Hibbett DS. 2010. Amylocorticiales ord. nov. and Jaapiales ord. nov.: early diverging clades of Agaricomycetidae dominated by corticioid forms. Mycologia 102(4):865−880

doi: 10.3852/09-288

[152]

He ZM, Yang ZL. 2022. The genera Bonomyces, Harmajaea and Notholepista from Northwestern China: two new species and a new record. Mycological Progress 21(2):26

doi: 10.1007/s11557-022-01786-0

[153]

He ZM, Yang ZL. 2023. A contribution to the knowledge of the genus Infundibulicybe (Tricholomatineae, Agaricales) in China: two new species and five redescribed taxa. Mycologia 115(5):693−713

doi: 10.1080/00275514.2023.2226893

[154]

Wijesinghe SN, Deng Y, Yuan Q, Zhou H, Wang L, et al. 2025. Mycosphere Notes 572–624: exploring the hidden diversity of fungi and fungilike taxa in different terrestrial microhabitats. Mycosphere 16(1):2975−3129

doi: 10.5943/mycosphere/16/1/21

[155]

Hu JJ, Tuo YL, Qi ZX, Jiang DH, Li Y, Zhang B. 2025. Two new species of Gymnopus sect. Levipedes (Omphalotaceae, Agaricales) from Central and North China. MycoKeys 118:19−34

doi: 10.3897/mycokeys.118.143277

[156]

Li JP, Gao J, Li Y, Xie ML, Li CT. 2026. New additions to Omphalotaceae (Agaricales, Basidiomycota): two new rhizomorph-forming species from China. MycoKeys 131:41−59

doi: 10.3897/mycokeys.131.184214

[157]

Deng CY, Wang WK, Li JP, Antonín V. 2025. Morphology and phylogeny reveal Gymnopus pseudoandrosaceus sp. nov. (Agaricales, Basidiomycota) from southwest China. Phytotaxa 683(3):218−228

doi: 10.11646/phytotaxa.683.3.3

[158]

Feng WC, Li XY, Cui YY, Cai Q. 2025. Species diversity of Gymnopus section Levipedes in southwestern China, with a description of three new species. Journal of Fungi 11(2):88

doi: 10.3390/jof11020088

[159]

Fan LF, Wang BY, Ma TF, Li B, Ma JW, et al. 2024. Three new species of Gymnopus and Mycena (Agaricales, Basidiomycota) from Northwestern China. Frontiers in Microbiology 15:1487598

doi: 10.3389/fmicb.2024.1487598

[160]

Manawasinghe IS, Hyde KD, Wanasinghe DN, Karunarathna SC, Maharachchikumbura S, et al. 2025. Fungal diversity notes 1818–1918: taxonomic and phylogenetic contributions on genera and species of fungi. Fungal Diversity 130(1):1−261

doi: 10.1007/s13225-024-00541-y

[161]

Hu JJ, Tuo YL, Qi ZX, Li XF, Jiang DH, et al. 2024. The combination of morphological and phylogenetic evidence reveals four new Gymnopus species and new distribution. Journal of Fungi 10(10):672

doi: 10.3390/jof10100672

[162]

Mata JL, Hughes KW, Petersen R. 2006. An investigation of /omphalotaceae (Fungi: Euagarics) with emphasis on the genus Gymnopus. Sydowia 58:191−289

[163]

Hu JJ, Zhang B. 2023. 广义金钱菌属真菌研究概况[Overview of research on fungi of the genus Flococcus in a broad sense]. 菌物学报[Acta Mycologica Sinica] 42(1):143−159 (in Chinese)

doi: 10.13346/j.mycosystema.220419

[164]

Oliveira JJS, Vargas-Isla R, Cabral TS, Rodrigues DP, Ishikawa NK. 2019. Progress on the phylogeny of the Omphalotaceae: Gymnopus s. str., Marasmiellus s. str., Paragymnopus gen. nov. and Pusillomyces gen. nov. Mycological Progress 18(5):713−739

doi: 10.1007/s11557-019-01483-5

[165]

Noordeloos ME, Antonín V. 2008. Contribution to a monograph of marasmioid and collybioid fungi in Europe. Czech Mycology 60:21−27

doi: 10.33585/cmy.60103

[166]

Fries E. 1821. Systema mycologicum : sistens fungorum ordines, genera et species, huc usque cognitas, quas ad normam methodi naturalis determinavit. Lundae: Ex Officina Berlingiana doi: 10.5962/bhl.title.5378

[167]

Antonín V, Halling RE, Noordeloos ME. 1997. Generic concepts within the groups of Marasmius and Collybia sensu lato. Mycotaxon 63:359−368

doi: 10.5962/p.415332

[168]

Antonín V, Ryoo R, Ka KH. 2014. Marasmioid and gymnopoid fungi of the Republic of Korea. 7. Gymnopus sect. Androsacei. Mycological Progress 13:703−718

doi: 10.1007/s11557-013-0953-z

[169]

Desjardin DE, Halling RE, Hemmes DE. 1999. Agaricales of the Hawaiian Islands. 5. The genera Rhodocollybia and Gymnopus. Mycologia 91:166−176

doi: 10.1080/00275514.1999.12061006

[170]

Sun YL, Luo Y, Bau T. 2021. 吉林省担子菌补记(十)[Notes on basidiomycetes of Jilin province (X)]. 菌物研究 [Journal of Fungal Research] 19:139−147 (in Chinese)

doi: 10.13341/j.jfr.2021.1432

[171]

Petersen RH, Hughes KW. 2021. Collybiopsis and its type species, Co. ramealis. Mycotaxon 136(2):263−349

doi: 10.5248/136.263

[172]

Ryoo R, Antonín V, Ka KH, Tomšovský M. 2016. Marasmioid and gymnopoid fungi of the Republic of Korea. 8. Gymnopus section Impudicae. Phytotaxa 286(2):75−88

doi: 10.11646/phytotaxa.286.2.2

[173]

Sesli E. 2022. Gymnopus impudicus (Fr.) Antonín, Halling & Noordel.: a new record for the Turkish mycota. Biological Diversity and Conservation 15:103−106

doi: 10.46309/biodicon.2022.1081652

[174]

Antonín V, Noordeloos ME. 2010. A monograph of marasmioid and collybioid fungi in Europe: with 131 figures and 130 coloured plates. Eching, Germany: IHW-Verlag

[175]

Antonín V, Sedlák P, Tomšovský M. 2013. Taxonomy and phylogeny of European Gymnopus subsection levipedes (Basidiomycota, Omphalotaceae). Persoonia – Molecular Phylogeny and Evolution of Fungi 31:179−187

doi: 10.3767/003158513X674043

[176]

The New York Botanical Garden. 1907. North American flora. New York: New York Botanical Garden

[177]

Li JP, Oliveira J, Pan MC, Deng CY, Antonín V, et al. 2024. Notes on all genera of Omphalotaceae: expanding the taxonomic spectrum in China and revisiting historical type specimens. Mycosphere 15(1):1522−1594

doi: 10.5943/mycosphere/15/1/13

[178]

Hu JJ, Song LR, Tuo YL, Zhao GP, Yue L, et al. 2022. Multiple evidences reveal new species and a new record of smelly Gymnopus (Agaricales, Omphalotaceae) from China. Frontiers in Microbiology 13:968617

doi: 10.3389/fmicb.2022.968617

[179]

Wang GS, Cai Q, Hao YJ, Bau T, Chen ZH, et al. 2023. Phylogenetic and taxonomic updates of Agaricales, with an emphasis on Tricholomopsis. Mycology 15(2):180−209

doi: 10.1080/21501203.2023.2263031

[180]

Thapa A, Tamang J, Acharya K. 2023. A new species of Tricholomopsis (Agaricales) from Darjeeling Hills, India. Italian Journal of Mycology 52:144−153

doi: 10.6092/issn.2531-7342/18381

[181]

Wang G, Guo T, Chen X, Liao J, Ma Y, et al. 2026. Two new species of Tricholomopsis (Agaricales) from China based on molecular phylogenetic and morphological evidence. MycoKeys 131:235−249

doi: 10.3897/mycokeys.131.185587

[182]

Hosen MI, Xu JY, Li T, Gates G, Li TH. 2020. Tricholomopsis rubroaurantiaca, a new species of Tricholomataceae from southern China. Mycoscience 61(6):342−347

doi: 10.1016/j.myc.2020.06.005

[183]

Holec J, Kunca V, Kolařík M. 2019. Tricholomopsis badinensis sp. nov. and T. sulphureoides—two rare fungi of European old-growth forests. Mycological Progress 18(3):321−334

doi: 10.1007/s11557-018-1449-7

[184]

Lu JR, Lei L, Yu FM, Zhu XT, Liang JF, et al. 2025. Diversity of macrofungi in southeast Xizang, China 2. Mycology 17(1):1−57

doi: 10.1080/21501203.2025.2525864

[185]

Smith AH. 1960. Tricholomopsis (Agaricales) in the Western hemisphere. Brittonia 12:41−70

doi: 10.2307/2805334

[186]

Choudhary S, Mehmood T, Sharma YP, Uniyal P. 2025. Two novel species of genus Melanoleuca (Agaricales) from Indian Himalaya. Feddes Repertorium 136(4):277−286

doi: 10.1002/fedr.202400027

[187]

Antonín V, Ďuriška O, Jančovičová S, Kudláček T, Para R, et al. 2022. Melanoleuca monticola and M. romanensis, two new European species of Melanoleuca, and comments to M. graminicola group. Preprint

[188]

Voto P, Maraia G, Milanese V. 2021. Melanoleuca chalcibasis sp. nov. from the Bosco Nordio (Italy). Mycological Observations 1:35−43

[189]

Qi Y, Li CH, Song YM, Zhang M, Guo HB, et al. 2022. Melanoleuca subgriseoflava and M. substridula—two new Melanoleuca species (Agaricales, Basidiomycota) described from China. PeerJ 10:e13807

doi: 10.7717/peerj.13807

[190]

Antonín V, Ševčíková H, Para R, Ďuriška O, Kudláček T, et al. 2021. Melanoleuca galbuserae, M. fontenlae and M. acystidiata—three new species in subgenus Urticocystis (Pluteaceae, Basidiomycota) with Comments on M. castaneofusca and related species. Journal of Fungi 7(3):191

doi: 10.3390/jof7030191

[191]

Pei Y, Guo HB, Liu TZ, Qin WQ, Zhao D, et al. 2021. Three new Melanoleuca species (Agaricales, Basidiomycota) from north-eastern China, supported by morphological and molecular data. MycoKeys 80:133−148

doi: 10.3897/mycokeys.80.64369

[192]

Armada F. 2020. Quelques basidiomycètes nouveaux ou rares pour l'Andalousie, découverts près de Grenade dans la Sierra Nevada (Espagne). Bulletin Mycologique et Botanique Dauphiné-Savoie 239:67−78

[193]

Crous PW, Cowan DA, Maggs-Kölling G, Yilmaz N, Larsson E, et al. 2020. Fungal Planet description sheets: 1112−1181. Persoonia – Molecular Phylogeny and Evolution of Fungi 45:251−409

doi: 10.3767/persoonia.2020.45.10

[194]

Ullah Z, Khurshid R, Khan MB, Ahmad I, Jabeen S, et al. 2020. Melanoleuca kashmirensis sp. nov. in subg. Urticocystis from Pakistan. Phytotaxa 434(1):89−100

doi: 10.11646/phytotaxa.434.1.6

[195]

Vizzini A, Para R, Fontenla R, Ghignone S, Ercole E. 2012. A preliminary ITS phylogeny of Melanoleuca (Agaricales), with special reference to European taxa. Mycotaxon 118(1):361−381

doi: 10.5248/118.361

[196]

Boekhout T. 1988. Notulae ad floram agaricinam neerlandicam—XVI. New taxa, new combinations in Melanoleuca Pat. and notes on rare species in the Netherlands. Persoonia – Molecular Phylogeny and Evolution of Fungi 13(4):397−431

[197]

Antonín V, Ďuriška O, Jančovičová S, Kudláček T, Para R, et al. 2023. Two new European species of Melanoleuca (Fungi, Agaricales) with comments on the M. graminicola group. Systematics and Biodiversity 21:2218375

doi: 10.1080/14772000.2023.2218375

[198]

Yu XD, Lv SX, Ma D, Li FF, Lin Y, et al. 2014. Two new species of Melanoleuca (Agaricales, Basidiomycota) from northeastern China, supported by morphological and molecular data. Mycoscience 55(6):456−461

doi: 10.1016/j.myc.2014.01.007

[199]

Elahi H, Niazi AR. 2024. Melanoleuca lalsuhanraensis, a new species from Lal Suhanra National Park, Punjab, Pakistan. Nordic Journal of Botany 7:e04245

doi: 10.1111/njb.04245

[200]

Bigelow HE. 1973. The genus Clitocybula. Mycologia 65(5):1101−1116

doi: 10.1080/00275514.1973.12019530

[201]

Antonín V, Borovička J, Holec J, Piltaver A, Kolařík M. 2019. Taxonomic update of Clitocybula sensu lato with a new generic classification. Fungal biology 123(6):431−447

doi: 10.1016/j.funbio.2019.03.004

[202]

Santamaría N, Rubio Casas L, Zamora JC. 2022. Clitocybula ellipsospora, a new species found in the Iberian Peninsula. Fungi Iberici 2:75−88

doi: 10.51436/funiber/02.005

[203]

Na Q, Zeng H, Hu Y, Ding H, Ke B, et al. 2024. Morphological and phylogenetic analyses reveal five new species of Porotheleaceae (Agaricales, Basidiomycota) from China. MycoKeys 105:49−95

doi: 10.3897/mycokeys.105.118826

[204]

Singer R. 1978. Keys for the identification of the species of Agaricales II. Sydowia 31(1–6):193−237

[205]

Malysheva E, Morozova O, Contu M. 2011. New combinations in Clitocybula: a study of cystidiate Pseudoomphalina species (Basidiomycota, Agaricomycetes). Sydowia 63(1):85−104

[206]

Antonín V, Beran M, Borovička J, Dvořák D, Holec J. 2011. Clitocybula familia (Fungi, Agaricales)—taxonomy, distribution, ecology and first records in the Czech Republic and Slovakia. Czech Mycology 63(1):1−11

doi: 10.33585/cmy.63101

[207]

Ding XX, Xu X, Cui YY, Kost G, Wang PM, et al. 2023. A fifty-locus phylogenetic analysis provides deep insights into the phylogeny of Tricholoma (Tricholomataceae, Agaricales). Persoonia - Molecular Phylogeny and Evolution of Fungi 50:1−26

doi: 10.3767/persoonia.2023.50.01

[208]

Tedersoo L, May TW, Smith ME. 2010. Ectomycorrhizal lifestyle in fungi: global diversity, distribution, and evolution of phylogenetic lineages. Mycorrhiza 20(4):217−263

doi: 10.1007/s00572-009-0274-x

[209]

Mao N, Yang JJ, Zhang YX, Li T, Fan L. 2025. Species diversity of Tricholoma (Agaricales, Tricholomataceae) from Shanxi Province of northern China with the description of four new species. MycoKeys 112:59−80

doi: 10.3897/mycokeys.112.132652

[210]

Bessette AE, Bessette AR, Roody WC, Trudell SA. 2013. Tricholomas of North America: a mushroom field guide. Austin: University of Texas Press. pp. 1–220 doi: 10.7560/742338

[211]

Heilmann-Clausen J, Christensen M, Frøslev TG, Kjøller R. 2017. Taxonomy of Tricholoma in northern Europe based on ITS sequence data and morphological characters. Persoonia - Molecular Phylogeny and Evolution of Fungi 38:38−57

doi: 10.3767/003158517X693174

[212]

Reschke K, Popa F, Yang ZL, Kost G. 2018. Diversity and taxonomy of Tricholoma species from Yunnan, China, and notes on species from Europe and North America. Mycologia 110(6):1081−1109

doi: 10.1080/00275514.2018.1512295

[213]

Kost G. 1981. Vergleichende morphologische, anatomische und feinstrukturelle Merkmalsstudien an Arten der Gattung Tricholoma (Fr. ) Staude, Sektion Genuina (Fr. ) Sacc. PhD dissertation. Eberhard-Karls-Universität, Tübingen, Germany. pp. 1–152

[214]

Christensen M, Noordeloos ME. 1999. Notulae ad Floram agaricinam neerlandicam—XXXVI. Tricholoma. Persoonia–Molecular Phylogeny and Evolution of Fungi 17(Part 2):295−317

[215]

Christensen M, Heilmann-Clausen J. 2013. The genus Tricholoma. 1st ed. Denmark: Svampetryk. pp.1–228

[216]

Ding XX, Cui YY, Yang ZL. 2022. Two new species of Tricholoma sect. Genuina (Agaricales) from China based on molecular phylogenetic and morphological evidence. Mycological Progress 21(3):37

doi: 10.1007/s11557-022-01797-x

[217]

Huang HY, Wang R, Zhang WH, Yu FQ, Tang LP. 2024. Phylogeny and taxonomic revision of Hygrophorus robustus (Hygrophoraceae, Agaricales) from southwestern China. Phytotaxa 664(3):181−190

doi: 10.11646/phytotaxa.664.3.3

[218]

Yang JJ, Xu YY, Lv JC, Fan L. 2023. Tricholoma lishanense (Tricholomataceae, Agaricales), a new species from China revealed by morphology and phylogenetic analysis. Phytotaxa 598(1):89−96

doi: 10.11646/phytotaxa.598.1.7

[219]

Cui YY, Ding XX, Kost GW, Yang ZL. 2022. Tricholoma sect. Tricholoma (Tricholomataceae) from China: molecular phylogeny and taxonomy. Mycological Progress 21(3):35

doi: 10.1007/s11557-022-01788-y

[220]

Kaur M, Rather H, Malik N. 2021. Morphoanatomical characterization of an ectomycorrhizal species of Tricholoma (Tricholomataceae) from Kashmir Himalaya, India. Open Access Journal of Agricultural Research 6(1):16000262

doi: 10.23880/oajar-16000262

[221]

Ovrebo CL, Halling RE, Hughes KW, Kuo M. 2021. New species and records of Tricholoma (Agaricales: Tricholomataceae) sections Genuina and Megatricholoma from Costa Rica and United States. Journal of the Botanical Research Institute of Texas 15(2):545−557

doi: 10.17348/jbrit.v15.i2.1160

[222]

Põlme S, Abarenkov K, Henrik Nilsson R, Lindahl BD, Clemmensen KE, et al. 2020. FungalTraits: a user-friendly traits database of fungi and fungus-like stramenopiles. Fungal Diversity 105(1):1−16

doi: 10.1007/s13225-020-00466-2

[223]

Seitzman BH, Ouimette A, Mixon RL, Hobbie EA, Hibbett DS. 2011. Conservation of biotrophy in Hygrophoraceae inferred from combined stable isotope and phylogenetic analyses. Mycologia 103(2):280−290

doi: 10.3852/10-195

[224]

Song HB, Bau T. 2024. Resolving the polyphyletic origins of Pholiotina s.l. (Bolbitiaceae, Agaricales) based on Chinese materials and reliable foreign sequences. Mycosphere 15(1):1595−1674

doi: 10.5943/mycosphere/15/1/14

[225]

Dong J, Li Q, Yuan Q, Luo Y, Zhang X, et al. 2024. Species diversity, taxonomy, molecular systematics and divergence time of wood-inhabiting fungi in Yunnan-Guizhou Plateau, Asia. Mycosphere 15(1):1110−1293

doi: 10.5943/mycosphere/15/1/10

[226]

Guo MJ, Zhuo L, Wang S, Sui XN, Zhou H, et al. 2024. Hyperdiverse Rhytismatales on twigs of Rhododendron spp. Mycosphere 15(1):764−880

doi: 10.5943/mycosphere/15/1/6

[227]

Li JX, Zhao RL, Phurbu D, Xing R, Liu DM, et al. 2025. Disentangling taxonomic chaos in Agaricaceae s.l.: an integrative phylogenomic framework with divergence dating reconstructs classification. Fungal Diversity 135(1):745−881

doi: 10.1007/s13225-025-00568-9

[228]

Kumar S, Das R, Sharma P, Tamang S, Ranjan RK, et al. 2023. Evaluation of fungal diversity in high altitude soils in different temperature conditions. Journal of Climate Change 9(2):65−75

doi: 10.3233/jcc230016

[229]

Meng Q, Xie Z, Xu H, Guo J, Tang Y, et al. 2022. Out of the Qinghai-Tibetan plateau: origin, evolution and historical biogeography of Morchella (both Elata and Esculenta clades). Frontiers in Microbiology 13:1078663

doi: 10.3389/fmicb.2022.1078663

[230]

Loughnan D, Gilbert B. 2017. Trait-mediated community assembly: distinguishing the signatures of biotic and abiotic filters. Oikos 126(8):1112−1122

doi: 10.1111/oik.03945

[231]

Wang Y, Ma YH, Wu G, Yang XY, Liu YJ, et al. 2024. Polyphasic taxonomy clarifies the relationships between Butyriboletus and Exsudoporus, and new taxa and reports of Boletaceae from China. Mycosphere 15:881−953

doi: 10.5943/mycosphere/15/1/7

[232]

Zhou Q, Cai Y, Zhou L, Zhang H, Li J, et al. 2026. Strategic and diversified soil microbes: unraveling the belowground control of plant multidiversity-productivity relationships across elevation. CATENA 262:109613

doi: 10.1016/j.catena.2025.109613

[233]

Lin H, Wang X, Zhong M, Tang K, Du Y, et al. 2024. Elevational patterns and assembly processes of multifaceted bird diversity in a subtropical mountain system. Journal of Biogeography 51(7):1276−1289

doi: 10.1111/jbi.14827

[234]

Wang X, Zhong M, Yang S, Jiang J, Hu J. 2022. Multiple β-diversity patterns and the underlying mechanisms across amphibian communities along a subtropical elevational gradient. Diversity and Distributions 28(12):2489−2502

doi: 10.1111/ddi.13618

[235]

Wambulwa MC, Milne R, Wu ZY, Spicer RA, Provan J, et al. 2021. Spatiotemporal maintenance of flora in the Himalaya biodiversity hotspot: current knowledge and future perspectives. Ecology and Evolution 11(16):10794−10812

doi: 10.1002/ece3.7906

[236]

Liu F, Li JX, Yang WQ, He MQ, Cao B, et al. 2025. Evolution and horizontal transfer of the psilocybin biosynthetic gene cluster drive the diversification of magic mushrooms. Mycosphere 16(1):3673−3694

doi: 10.5943/mycosphere/16/1/28

[237]

Xie Y, MacKinnon J, Li D. 2004. Study on biogeographical divisions of China. Biodiversity & Conservation 13:1391−1417

doi: 10.1023/B:BIOC.0000019396.31168.ba

[238]

Päckert M, Favre A, Schnitzler J, Martens J, Sun YH, et al. 2020. "Into and Out of" the Qinghai-Tibet Plateau and the Himalayas: centers of origin and diversification across five clades of Eurasian montane and alpine passerine birds. Ecology and Evolution 10(17):9283−9300

doi: 10.1002/ece3.6615

[239]

Wen J, Zhang JQ, Nie ZL, Zhong Y, Sun H. 2014. Evolutionary diversifications of plants on the Qinghai-Tibetan Plateau. Frontiers in Genetics 5:4

doi: 10.3389/fgene.2014.00004

[240]

Rahbek C, Borregaard MK, Antonelli A, Colwell RK, Holt BG, et al. 2019. Building mountain biodiversity: geological and evolutionary processes. Science 365(6458):1114−1119

doi: 10.1126/science.aax0151

[241]

Tzedakis PC, Lawson IT, Frogley MR, Hewitt GM, Preece RC. 2002. Buffered tree population changes in a quaternary refugium: evolutionary implications. Science 297(5589):2044−2047

doi: 10.1126/science.1073083

[242]

Rahbek C, Borregaard MK, Colwell RK, Dalsgaard B, Holt BG, et al. 2019. Humboldt's enigma: what causes global patterns of mountain biodiversity? Science 365(6458):1108−1113

doi: 10.1126/science.aax0149

[243]

Kemppinen J, Lembrechts JJ, Van Meerbeek K, Carnicer J, Chardon NI, et al. 2024. Microclimate, an important part of ecology and biogeography. Global Ecology and Biogeography 33(6):e13834

doi: 10.1111/geb.13834

[244]

Sun H, Zhang J, Deng T, Boufford DE. 2017. Origins and evolution of plant diversity in the Hengduan Mountains, China. Plant Diversity 39(4):161−166

doi: 10.1016/j.pld.2017.09.004

[245]

Dai YC, Yu CJ, Wang HC. 2007. Polypores from eastern Xizang (Tibet), western China. Annales Botanici Fennici 44(2):135−145

[246]

Dai YC, Yang ZL, Cui BK, Wu G, Yuan HS, et al. 2021. 中国森林大型真菌重要类群多样性和系统学研究[Diversity and systematics of the important macrofungi in Chinese forests]. 菌物学报[Mycosystema] 40(4):770−805 (in Chinese)

doi: 10.13346/j.mycosystema.210036

[247]

Yuan Y, Chen JJ, Korhonen K, Martin F, Dai YC. 2021. An updated global species diversity and phylogeny in the forest pathogenic genus Heterobasidion (Basidiomycota, Russulales). Frontiers in Microbiology 11:596393

doi: 10.3389/fmicb.2020.596393

[248]

Zhang QY, Liu HG, Papp V, Zhou M, Dai YC, et al. 2023. New insights into the classification and evolution of Favolaschia (Agaricales, Basidiomycota) and its potential distribution, with descriptions of eight new species. Mycosphere 14(1):777−814

doi: 10.5943/mycosphere/14/1/10

[249]

Zhang X, Zhou HM, Tu ZH, Zeng GY, Dai YC. 2026. Morphological and phylogenetic evidence reveal two new Pseudohydnum (Auriculariales, Basidiomycota) species from Guangxi and Xizang, China. MycoKeys 131:251−264

doi: 10.3897/mycokeys.131.189424

[250]

Zhou M, Dai YC, Vlasák J, Liu HG, Yuan Y. 2023. Updated systematics of Trichaptum s.l. (Hymenochaetales, Basidiomycota). Mycosphere 14(1):815−917

doi: 10.5943/mycosphere/14/1/11

[251]

He MQ, Yang WQ, Phurbu D, Liu F, Li JX, et al. 2026. Systematic study of Panaeolus (Agaricales, Galeropsidaceae) sensu lato and psilocybin-producing traits of species from China. IMA Fungus 17:e167329

doi: 10.3897/imafungus.17.167329

[252]

Gu Y, Meng L, Wang Y, Wu Z, Pan Y, et al. 2025. Uncovering the role of solar radiation and water stress factors in constraining decadal intra-site spring phenology variability in diverse ecosystems across the Northern Hemisphere. New Phytologist 246(5):1986−2003

doi: 10.1111/nph.70104

[253]

Zhang H, Zhang Y, Miralles DG, Tai X, Koppa A, et al. 2026. Increase in plant reliance on past precipitation associated with greening and drying. Nature Ecology & Evolution 10(4):689−699

doi: 10.1038/s41559-026-02997-4

[254]

Liu M, Peñuelas J, Trugman AT, Vargas GG, Yang L, et al. 2025. Diverging responses of terrestrial ecosystems to water stress after disturbances. Nature Climate Change 15(1):73−79

doi: 10.1038/s41558-024-02191-z

[255]

Krah FS, Hagge J, Schreiber J, Brandl R, Müller J, et al. 2022. Fungal fruit body assemblages are tougher in harsh microclimates. Scientific Reports 12:1633

doi: 10.1038/s41598-022-05715-9

[256]

Halbwachs H, Brandl R, Bässler C. 2015. Spore wall traits of ectomycorrhizal and saprotrophic agarics may mirror their distinct lifestyles. Fungal Ecology 17:197−204

doi: 10.1016/j.funeco.2014.10.003

[257]

Liu JW, Zhao WQ, Ren ML, Liu YQ, Xu Y, et al. 2022. Contrasting elevational patterns and underlying drivers of stream bacteria and fungi at the regional scale on the Tibetan Plateau. FEMS Microbiology Ecology 98(6):fiac050

doi: 10.1093/femsec/fiac050

[258]

Guo M, Xia Z, He X, Wan S, Wang Y, et al. 2026. High-throughput sequencing reveals endophytic bacterial differentiation of common truffles (Tuber spp.) in China: diversity, biogeographical patterns, and fungal health implications. Microbiology Spectrum 14(2):e01866-25

doi: 10.1128/spectrum.01866-25

[259]

Kraft NJB, Adler PB, Godoy O, James EC, Fuller S, et al. 2015. Community assembly, coexistence and the environmental filtering metaphor. Functional Ecology 29(5):592−599

doi: 10.1111/1365-2435.12345

[260]

Dorjey K, Kumar S, Sharma YP. 2023. The adaptations of high-altitude mushrooms in the cold desert of Ladakh. In Environmental Change and Development in Ladakh, Indian Trans-Himalaya, eds. Humbert-Droz B, Dame J, Morup T. Advances in Asian Human-Environmental Research. Cham: Springer Nature Switzerland. pp. 95–110 doi: 10.1007/978-3-031-42494-6_7

[261]

Halbwachs H, Bässler C. 2015. Gone with the wind—a review on basidiospores of lamellate agarics. Mycosphere 6(1):78−112

doi: 10.5943/mycosphere/6/1/10

[262]

Linder DH. 1940. Evolution of the basidiomycetes and its relation to the terminology of the basidium. Mycologia 32(4):419−447

doi: 10.1080/00275514.1940.12017422

[263]

Sonnenberg ASM, Sedaghat-Telgerd N, Lavrijssen B, Hendrickx PM, Scholtmeijer K, et al. 2021. Mapping recombination landscape and basidial spore number in the button mushroom Agaricus bisporus. Frontiers in Fungal Biology 2:711330

doi: 10.3389/ffunb.2021.711330

[264]

Pehim Limbu S, Stürmer SL, Zahn G, Aguilar-Trigueros CA, Rogers N, et al. 2025. Climate-linked biogeography of mycorrhizal fungal spore traits. Proceedings of the National Academy of Sciences U. S. A. 122(29):e2505059122

doi: 10.1073/pnas.2505059122

[265]

Meerts. 1999. The evolution of spore size in Agarics: do big mushrooms have big spores? Journal of Evolutionary Biology 12(1):161−165

doi: 10.1046/j.1420-9101.1999.00020.x

[266]

Crandall SG, Saarman N, Gilbert GS. 2020. Fungal spore diversity, community structure, and traits across a vegetation mosaic. Fungal Ecology 45:100920

doi: 10.1016/j.funeco.2020.100920

[267]

Zanne AE, Abarenkov K, Afkhami ME, Aguilar-Trigueros CA, Bates S, et al. 2020. Fungal functional ecology: bringing a trait-based approach to plant-associated fungi. Biological Reviews 95(2):409−433

doi: 10.1111/brv.12570

[268]

Dawson SK, Boddy L, Halbwachs H, Bässler C, Andrew C, et al. 2019. Handbook for the measurement of macrofungal functional traits: a start with basidiomycete wood fungi. Functional Ecology 33(3):372−387

doi: 10.1111/1365-2435.13239

[269]

Wang XW, Varga T, Li QS, G. Nagy LG, Zhou LW. 2025. Complex evolutionary history of the fungal order Hymenochaetales revealed by analyses of trait evolution and diversification. Mycosphere 16(1):517−535

doi: 10.5943/mycosphere/16/1/7

[270]

Zhao H, Cui YJ, Guan QX, Wang K, Zhuang L, et al. 2025. Global fungal diversity and distribution patterns within the order Hymenochaetales (Agaricomycetes, Basidiomycota). Mycosphere 16(1):3257−3280

doi: 10.5943/mycosphere/16/1/24

[271]

Cao GL, Li XQ, Xiang KL, Erst AS, Jabbour F, et al. 2025. Formation of three great Asian plateaus, climate change, and biodiversity. Trends in Ecology & Evolution 40(10):970−982

doi: 10.1016/j.tree.2025.07.008

[272]

Chen YH, Lenoir J, Chen IC. 2025. Limited evidence for range shift-driven extinction in mountain biota. Science 388(6748):741−747

doi: 10.1126/science.adq9512

[273]

Runnel K, Tedersoo L, Krah FS, Piepenbring M, Scheepens JF, et al. 2025. Toward harnessing biodiversity-ecosystem function relationships in fungi. Trends in Ecology & Evolution 40(2):180−190

doi: 10.1016/j.tree.2024.10.004

[274]

Sun Y, Deng Y, Yao S, Sun Y, Degen AA, et al. 2025. Distribution range and richness of plant species are predicted to increase by 2100 due to a warmer and wetter climate in northern China. Global Change Biology 31(7):e70334

doi: 10.1111/gcb.70334

[275]

Pecl GT, Araújo MB, Bell JD, Blanchard J, Bonebrake TC, et al. 2017. Biodiversity redistribution under climate change: impacts on ecosystems and human well-being. Science 355(6332):eaai9214

doi: 10.1126/science.aai9214

[276]

Han XX, Cao B, He MQ, Zhu XY, Li JX, et al. 2024. 中国西南地区沙鲁里山系大型真菌物种多样性[Species diversity of macrofungi in Shaluli Mountains, southwest China]. 菌物学报[MYCOSYSTEMA] 43(3):206−219 (in Chinese)

doi: 10.13346/j.mycosystema.230304

[277]

Liu DM, Wang SH, Wang K, Li JX, Yang WQ, et al. 2025. Species diversity and resource status of macrofungi in Beijing: insights from natural and urban habitats. Journal of Fungi 11(8):607

doi: 10.3390/jof11080607

[278]

Glamočlija J, Stojković D, Nikolić M, Ćirić A, Reis FS, et al. 2015. A comparative study on edible Agaricus mushrooms as functional foods. Food & function 6(6):1900−1910

doi: 10.1039/C4FO01135J

[279]

He MQ, Wang MQ, Chen ZH, Deng WQ, Li TH, et al. 2022. Potential benefits and harms: a review of poisonous mushrooms in the world. Fungal Biology Reviews 42:56−68

doi: 10.1016/j.fbr.2022.06.002

[280]

Shi J, Wang X, Wang E. 2023. Mycorrhizal symbiosis in plant growth and stress adaptation: from genes to ecosystems. Annual review of plant biology 74:569−607

doi: 10.1146/annurev-arplant-061722-090342

[281]

Martin F, Tan H. 2025. Saprotrophy-to-symbiosis continuum in fungi. Current Biology 35(11):R469−R475

doi: 10.1016/j.cub.2025.01.032