[1]

Masson-Delmotte V, Zhai P, Pirani A, Connors SL, Péan C, et al. 2021. Climate change 2021: the physical science basis. Contribution of working group I to the sixth assessment report of the intergovernmental panel on climate change. Cambridge, United Kingdom: Cambridge University Press. doi: 10.1017/9781009157896

[2]

Kang Y, Khan S, Ma X. 2009. Climate change impacts on crop yield, crop water productivity and food security – a review. Progress in Natural Science 19(12):1665−1674

doi: 10.1016/j.pnsc.2009.08.001
[3]

Lobell DB, Gourdji SM. 2012. The influence of climate change on global crop productivity. Plant Physiology 160(4):1686−1697

doi: 10.1104/pp.112.208298
[4]

Song HJ, Mishra U, Park SY, Seo YH, Turner BL, et al. 2025. Warming but not elevated CO2 depletes soil organic carbon in a temperate rice paddy. Agriculture, Ecosystems & Environment 379:109333

doi: 10.1016/j.agee.2024.109333
[5]

Tang S, Cheng W, Hu R, Guigue J, Hattori S, et al. 2021. Five-year soil warming changes soil C and N dynamics in a single rice paddy field in Japan. Science of The Total Environment 756:143845

doi: 10.1016/j.scitotenv.2020.143845
[6]

United Nations Department for Economic and Social Affairs. 2025. World population prospects 2024: summary of results. New York, NY: United Nations

[7]

Fischer T, Byerlee D, Edmeades G. 2014. Crop yields and global food security: will yield increase continue to feed the world? Canberra, Australia: ACIAR

[8]

Liu X, Lie Z, Reich PB, Zhou G, Yan J, et al. 2024. Long-term warming increased carbon sequestration capacity in a humid subtropical forest. Global Change Biology 30(1):e17072

doi: 10.1111/gcb.17072
[9]

Hicks Pries CE, Castanha C, Porras RC, Torn MS. 2017. The whole-soil carbon flux in response to warming. Science 355(6332):1420−1423

doi: 10.1126/science.aal1319
[10]

Soong JL, Castanha C, Hicks Pries CE, Ofiti N, Porras RC, et al. 2021. Five years of whole-soil warming led to loss of subsoil carbon stocks and increased CO 2 efflux. Science Advances 7(21):eabd1343

doi: 10.1126/sciadv.abd1343
[11]

Zhang G, Kang Y, Han G, Sakurai K. 2011. Effect of climate change over the past half century on the distribution, extent and NPP of ecosystems of Inner Mongolia: effect of climate change on the ecosystem. Global Change Biology 17(1):377−389

doi: 10.1111/j.1365-2486.2010.02237.x
[12]

Lie Z, Huang W, Liu X, Zhou G, Yan J, et al. 2021. Warming leads to more closed nitrogen cycling in nitrogen-rich tropical forests. Global Change Biology 27(3):664−674

doi: 10.1111/gcb.15432
[13]

Hall SJ, Tenesaca CG, Lawrence NC, Green DIS, Helmers MJ, et al. 2023. Poorly drained depressions can be hotspots of nutrient leaching from agricultural soils. Journal of Environmental Quality 52(3):678−690

doi: 10.1002/jeq2.20461
[14]

Li Y, Zou N, Liang X, Zhou X, Guo S, et al. 2023. Effects of nitrogen input on soil bacterial community structure and soil nitrogen cycling in the rhizosphere soil of Lycium barbarum L. Frontiers in Microbiology 13:1070817

doi: 10.3389/fmicb.2022.1070817
[15]

Nelson MB, Martiny AC, Martiny JBH. 2016. Global biogeography of microbial nitrogen-cycling traits in soil. Proceedings of the National Academy of Sciences of the United States of America 113(29):8033−8040

doi: 10.1073/pnas.1601070113
[16]

Van Groenigen JW, Huygens D, Boeckx P, Kuyper TW, Lubbers IM, et al. 2015. The soil N cycle: new insights and key challenges. Soil 1(1):235−256

doi: 10.5194/soil-1-235-2015
[17]

Zhu X, Zhang W, Chen H, Mo J. 2015. Impacts of nitrogen deposition on soil nitrogen cycle in forest ecosystems: a review. Acta Ecologica Sinica 35(3):35−43

doi: 10.1016/j.chnaes.2015.04.004
[18]

Salo T, Turtola E. 2006. Nitrogen balance as an indicator of nitrogen leaching in Finland. Agriculture, Ecosystems & Environment 113(1):98−107

doi: 10.1016/j.agee.2005.09.002
[19]

Shi Z, Li D, Jing Q, Cai J, Jiang D, et al. 2012. Effects of nitrogen applications on soil nitrogen balance and nitrogen utilization of winter wheat in a rice–wheat rotation. Field Crops Research 127:241−247

doi: 10.1016/j.fcr.2011.11.025
[20]

Jiang D, Chen L, Xia N, Norgbey E, Koomson DA, et al. 2020. Elevated atmospheric CO2 impact on carbon and nitrogen transformations and microbial community in replicated wetland. Ecological Processes 9(1):57

doi: 10.1186/s13717-020-00267-0
[21]

Fanin N, Mooshammer M, Sauvadet M, Meng C, Alvarez G, et al. 2022. Soil enzymes in response to climate warming: mechanisms and feedbacks. Functional Ecology 36(6):1378−1395

doi: 10.1111/1365-2435.14027
[22]

Li J, Wang G, Mayes MA, Allison SD, Frey SD, et al. 2019. Reduced carbon use efficiency and increased microbial turnover with soil warming. Global Change Biology 25(3):900−910

doi: 10.1111/gcb.14517
[23]

Hagerty SB, Van Groenigen KJ, Allison SD, Hungate BA, Schwartz E, et al. 2014. Accelerated microbial turnover but constant growth efficiency with warming in soil. Nature Climate Change 4(10):903−906

doi: 10.1038/nclimate2361
[24]

Bai E, Li S, Xu W, Li W, Dai W, et al. 2013. A meta-analysis of experimental warming effects on terrestrial nitrogen pools and dynamics. New Phytologist 199(2):441−451

doi: 10.1111/nph.12252
[25]

Gu J, Yang J. 2022. Nitrogen (N) transformation in paddy rice field: its effect on N uptake and relation to improved N management. Crop and Environment 1(1):7−14

doi: 10.1016/j.crope.2022.03.003
[26]

Rural Development Administration (RDA). 2017. Fertilization standard of crop plants. RDA, South Korea

[27]

Rolston DE. 1986. Gas flux. In Methods of Soil Analysis, Part 1, Second edition, ed. Madison KA, WI: ASA and SSSA. pp. 1103–1119 doi: 10.2136/sssabookser5.1.2ed.c47

[28]

Singh S, Singh JS, Kashyap AK. 1999. Methane flux from irrigated rice fields in relation to crop growth and N-fertilization. Soil Biology and Biochemistry 31(9):1219−1228

doi: 10.1016/S0038-0717(99)00027-9
[29]

Christensen BT. 2001. Physical fractionation of soil and structural and functional complexity in organic matter turnover. European Journal of Soil Science 52(3):345−353

doi: 10.1046/j.1365-2389.2001.00417.x
[30]

Cotrufo MF, Soong JL, Horton AJ, Campbell EE, Haddix ML, et al. 2015. Formation of soil organic matter via biochemical and physical pathways of litter mass loss. Nature Geoscience 8(10):776−779

doi: 10.1038/ngeo2520
[31]

Lavallee JM, Soong JL, Cotrufo MF. 2020. Conceptualizing soil organic matter into particulate and mineral-associated forms to address global change in the 21st century. Global Change Biology 26(1):261−273

doi: 10.1111/gcb.14859
[32]

Zehr JP, Jenkins BD, Short SM, Steward GF. 2003. Nitrogenase gene diversity and microbial community structure: a cross-system comparison. Environmental Microbiology 5:539−554

doi: 10.1046/j.1462-2920.2003.00451.x
[33]

Poly F, Monrozier LJ, Bally R. 2001. Improvement in the RFLP procedure for studying the diversity of nifH genes in communities of nitrogen fixers in soil. Research in Microbiology 152(1):95−103

doi: 10.1016/S0923-2508(00)01172-4
[34]

Ritalahti KM, Amos BK, Sung Y, Wu Q, Koenigsberg SS, et al. 2006. Quantitative PCR targeting 16S rRNA and reductive dehalogenase genes simultaneously monitors multiple Dehalococcoides strains. Applied and Environmental Microbiology 72(4):2765−2774

doi: 10.1128/AEM.72.4.2765-2774.2006
[35]

Kim HY, Lieffering M, Kobayashi K, Okada M, Miura S. 2003. Seasonal changes in the effects of elevated CO 2 on rice at three levels of nitrogen supply: a free air CO2 enrichment (FACE) experiment. Global Change Biology 9(6):826−837

doi: 10.1046/j.1365-2486.2003.00641.x
[36]

Usui Y, Sakai H, Tokida T, Nakamura H, Nakagawa H, et al. 2016. Rice grain yield and quality responses to free-air CO2 enrichment combined with soil and water warming. Global Change Biology 22(3):1256−1270

doi: 10.1111/gcb.13128
[37]

Kirschbaum MUF. 2004. Direct and indirect climate change effects on photosynthesis and transpiration. Plant Biology 6(3):242−253

doi: 10.1055/s-2004-820883
[38]

Yoshimoto M, Oue H, Takahashi N, Kobayashi K. 2005. The effects of FACE (free-air CO2 enrichment) on temperatures and transpiration of rice panicles at flowering stage. Journal of Agricultural Meteorology 60(5):597−600

doi: 10.2480/agrmet.597
[39]

Robinson D. 2001. δ15N as an integrator of the nitrogen cycle. Trends in Ecology & Evolution 16(3):153−162

doi: 10.1016/S0169-5347(00)02098-X
[40]

Högberg P. 1997. Tansley review No. 95. 15N natural abundance in soil-plant systems. New Phytologist 137:179-203

doi: 10.1046/j.1469-8137.1997.00808.x
[41]

Buckeridge KM, Grogan P. 2008. Deepened snow alters soil microbial nutrient limitations in Arctic birch hummock tundra. Applied Soil Ecology 39(2):210−222

doi: 10.1016/j.apsoil.2007.12.010
[42]

Denk TRA, Mohn J, Decock C, Lewicka-Szczebak D, Harris E, et al. 2017. The nitrogen cycle: a review of isotope effects and isotope modeling approaches. Soil Biology and Biochemistry 105:121−137

doi: 10.1016/j.soilbio.2016.11.015
[43]

Liang C, Schimel JP, Jastrow JD. 2017. The importance of anabolism in microbial control over soil carbon storage. Nature Microbiology 2(8):17105

doi: 10.1038/nmicrobiol.2017.105
[44]

Jilling A, Grandy AS, Daly AB, Hestrin R, Possinger A, et al. 2025. Evidence for the existence and ecological relevance of fast-cycling mineral-associated organic matter. Communications Earth & Environment 6(1):690

doi: 10.1038/s43247-025-02681-8
[45]

Wild B, Alaei S, Bengtson P, Bodé S, Boeckx P, et al. 2017. Short-term carbon input increases microbial nitrogen demand, but not microbial nitrogen mining, in a set of boreal forest soils. Biogeochemistry 136(3):261−278

doi: 10.1007/s10533-017-0391-0
[46]

Finzi AC, Norby RJ, Calfapietra C, Gallet-Budynek A, Gielen B, et al. 2007. Increases in nitrogen uptake rather than nitrogen-use efficiency support higher rates of temperate forest productivity under elevated CO2. Proceedings of the National Academy of Sciences of the United States of America 104(35):14014−14019

doi: 10.1073/pnas.0706518104