| [1] |
Wang K, Liu H, Mei Q, Yang J, Ma F, et al. 2023. Characteristics of bHLH transcription factors and their roles in the abiotic stress responses of horticultural crops. |
| [2] |
Massari ME, Murre C. 2000. Helix-loop-helix proteins: regulators of transcription in eucaryotic organisms. |
| [3] |
Zhang C, Jiang L, Qian J, Yu G, Qing H, et al. 2025. Genome-wide analysis of basic helix-loop-helix (bHLH) transcription factors in petunia and identification of the putative candidate member involved in floral volatile benzenoids/phenylpropanoids metabolism. |
| [4] |
Mao K, Dong Q, Li C, Liu C, Ma F. 2017. Genome wide identification and characterization of apple bHLH transcription factors and expression analysis in response to drought and salt stress. |
| [5] |
Toledo-Ortiz G, Huq E, Quail PH. 2003. The Arabidopsis basic/helix-loop-helix transcription factor family. |
| [6] |
Feller A, Machemer K, Braun EL, Grotewold E. 2011. Evolutionary and comparative analysis of MYB and bHLH plant transcription factors. |
| [7] |
Kavas M, Baloğlu MC, Atabay ES, Ziplar UT, Daşgan HY, et al. 2016. Genome-wide characterization and expression analysis of common bean bHLH transcription factors in response to excess salt concentration. |
| [8] |
Zhao Q, Ren Y, Wang Q, Yao Y, You C, et al. 2016. Overexpression of MdbHLH104 gene enhances the tolerance to iron deficiency in apple. |
| [9] |
Huq E, Quail PH. 2002. PIF4, a phytochrome-interacting bHLH factor, functions as a negative regulator of phytochrome B signaling in Arabidopsis. |
| [10] |
Jiang Y, Yang B, Deyholos MK. 2009. Functional characterization of the Arabidopsis bHLH92 transcription factor in abiotic stress. |
| [11] |
Qi T, Wang J, Huang H, Liu B, Gao H, et al. 2015. Regulation of jasmonate-induced leaf senescence by antagonism between bHLH subgroup IIIe and IIId factors in arabidopsis. |
| [12] |
Goossens J, Mertens J, Goossens A. 2017. Role and functioning of bHLH transcription factors in jasmonate signalling. |
| [13] |
Khan RA, Abbas N. 2023. Role of epigenetic and post-translational modifications in anthocyanin biosynthesis: a review. |
| [14] |
Xie X, Li S, Zhang R, Zhao J, Chen Y, et al. 2012. The bHLH transcription factor MdbHLH3 promotes anthocyanin accumulation and fruit colouration in response to low temperature in apples. |
| [15] |
Lim SH, Kim DH, Jung JA, Lee JY. 2021. Alternative splicing of the basic helix-loop-helix transcription factor gene CmbHLH2 affects anthocyanin biosynthesis in ray florets of Chrysanthemum (Chrysanthemum morifolium). |
| [16] |
Zhao W, Liu Y, Li L, Meng H, Yang Y, et al. 2021. Genome-wide identification and characterization of bHLH transcription factors related to anthocyanin biosynthesis in red walnut (Juglans regia L.). |
| [17] |
Lepiniec L, Debeaujon I, Routaboul JM, Baudry A, Pourcel L, et al. 2006. Genetics and biochemistry of seed flavonoids. |
| [18] |
Baudry A, Heim MA, Dubreucq B, Caboche M, Weisshaar B, et al. 2004. TT2, TT8, and TTG1 synergistically specify the expression of BANYULS and proanthocyanidin biosynthesis in Arabidopsis thaliana. |
| [19] |
Li M, Sun L, Gu H, Cheng D, Guo X, et al. 2021. Genome-wide characterization and analysis of bHLH transcription factors related to anthocyanin biosynthesis in spine grapes (Vitis davidii). |
| [20] |
Ji X, Yang F, Zhou X, Jia W, Zhu X, et al. 2025. Genome-wide identification of the bHLH gene family and the mechanism regulation of anthocyanin biosynthesis by ChEGL1 in Cerasus humilis. |
| [21] |
Wang R, Mao C, Ming F. 2022. PeMYB4L interacts with PeMYC4 to regulate anthocyanin biosynthesis in Phalaenopsis orchid. |
| [22] |
Liu H, Liu Q, Chen Y, Zhu Y, Zhou X, et al. 2023. Full-length transcriptome sequencing provides insights into flavonoid biosynthesis in Camellia nitidissima Petals. |
| [23] |
Chase MW, Cameron KM, Freudenstein JV, Pridgeon AM, Salazar G, et al. 2015. An updated classification of Orchidaceae. |
| [24] |
Govaerts R, Campacci MA, Holland A, Pridgeon AM. 2024. World Checklist of Orchidaceae. Royal Botanic Gardens, Kew. http://apps.kew.org/wcsp/. |
| [25] |
Yang F, Gao J, Wei Y, Ren R, Zhang G, et al. 2021. The genome of Cymbidium sinense revealed the evolution of orchid traits. |
| [26] |
Wang Y, Li S, Zhang Q, Shi Z, Zhai G, et al. 2026. Genome-wide analysis of MYB transcription factors in Cymbidium species reveals their roles in anthocyanin synthesis. |
| [27] |
Gao F, Dubos C. 2024. The Arabidopsis bHLH transcription factor family. |
| [28] |
Wu Y, Wu S, Wang X, Mao T, Bao M, et al. 2022. Genome-wide identification and characterization of the bHLH gene family in an ornamental woody plant Prunus mume. |
| [29] |
Chen C, Wu Y, Li J, Wang X, Zeng Z, et al. 2023. TBtools-II: a 'one for all, all for one' bioinformatics platform for biological big-data mining. |
| [30] |
Kumar S, Stecher G, Suleski M, Sanderford M, Sharma S, et al. 2024. MEGA12: molecular evolutionary genetic analysis version 12 for adaptive and green computing. |
| [31] |
Waterhouse AM, Procter JB, Martin DM, Clamp M, Barton GJ. 2009. Jalview Version 2—a multiple sequence alignment editor and analysis workbench. |
| [32] |
Zhu M, Wang Q, Tu S, Ke S, Bi Y, et al. 2023. Genome-wide identification analysis of the R2R3-MYB transcription factor family in Cymbidium sinense for insights into drought stress responses. |
| [33] |
Lalitha S. 2000. Primer premier 5. Biotech Software & Internet Report 1:270−272 |
| [34] |
Livak KJ, Schmittgen TD. 2001. Analysis of relative gene expression data using real-time quantitative PCR and the 2−ΔΔCᴛ Method. |
| [35] |
Carretero-Paulet L, Galstyan A, Roig-Villanova I, Martínez-García JF, Bilbao-Castro JR, et al. 2010. Genome-wide classification and evolutionary analysis of the bHLH family of transcription factors in Arabidopsis, poplar, rice, moss, and algae. |
| [36] |
Wang M, Ou Y, Li Z, Zheng Q, Ke Y, et al. 2023. Genome-wide identification and analysis of bHLH transcription factors related to anthocyanin biosynthesis in Cymbidium ensifolium. |
| [37] |
Zhai X, Wang X, Yang X, Huang Q, Wu D, et al. 2024. Genome-wide identification of bHLH transcription factors and expression analysis under drought stress in Pseudoroegneria libanotica at germination. |
| [38] |
Yang T, Zhu R, Li J, Wang Y, Zhou L, et al. 2026. A bHLH transcription factor negatively regulates effective panicle number and grain yield by modulating auxin transport and distribution in rice. |
| [39] |
Shen J, Su T, Yu D, Tan S, Zhang Y. 2025. Evolution by gene duplication: in the era of genomics. |
| [40] |
Gong Y, Chen X, Qian Z, Pan P, Sun Z, et al. 2026. Evolutionary and functional divergence of MYB regulatory networks in flavonoid biosynthesis across Poaceae, non-Poaceae monocots, and eudicot horticultural plants. |
| [41] |
Ludwig SR, Habera LF, Dellaporta SL, Wessler SR. 1989. Lc, a member of the maize R gene family responsible for tissue-specific anthocyanin production, encodes a protein similar to transcriptional activators and contains the myc-homology region. |
| [42] |
Albert NW, Davies KM, Lewis DH, Zhang H, Montefiori M, et al. 2014. A conserved network of transcriptional activators and repressors regulates anthocyanin pigmentation in eudicots. |
| [43] |
Hichri I, Heppel SC, Pillet J, Léon C, Czemmel S, et al. 2010. The basic helix-loop-helix transcription factor MYC1 is involved in the regulation of the flavonoid biosynthesis pathway in grapevine. |
| [44] |
Zhao K, Zhang Q, Wang Y, Wei Q, Wang Y. 2025. Advances in the biosynthesis and regulatory mechanisms of anthocyanins in horticultural plants: a comprehensive review. |