Taxonomic Studies of the Ground Beetle Subgenus Rhagadus Motschulsky (Coleoptera: Carabidae: Pterostichus): Unraveling the “Female Holotype Problem” Using External Geometric Morphometrics and Novel Male Genital Morphology

Kôji Sasakawa; Kôji Sasakawa

doi:10.1155/jzs/1776890

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Copyright: © 2025 by the author(s). Exclusive Licensee Maximum Academic Press, Fayetteville, GA. This article is an open access article distributed under Creative Commons Attribution License (CC BY 4.0), visit https://creativecommons.org/licenses/by/4.0/.

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Kôji Sasakawa. 2025. Taxonomic Studies of the Ground Beetle Subgenus Rhagadus Motschulsky (Coleoptera: Carabidae: Pterostichus): Unraveling the “Female Holotype Problem” Using External Geometric Morphometrics and Novel Male Genital Morphology. Journal of Zoological Systematics and Evolutionary Research. 2025: doi: 10.1155/jzs/1776890

Kôji Sasakawa. 2025. Taxonomic Studies of the Ground Beetle Subgenus Rhagadus Motschulsky (Coleoptera: Carabidae: Pterostichus): Unraveling the “Female Holotype Problem” Using External Geometric Morphometrics and Novel Male Genital Morphology. Journal of Zoological Systematics and Evolutionary Research. 2025: doi: 10.1155/jzs/1776890

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Taxonomic Studies of the Ground Beetle Subgenus Rhagadus Motschulsky (Coleoptera: Carabidae: Pterostichus): Unraveling the “Female Holotype Problem” Using External Geometric Morphometrics and Novel Male Genital Morphology

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